Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Panorpida
Amphiesmenoptera
Order Lepidoptera Linnaeus, 1758 - butterflies, moths
Suborder Glossata Fabricius, 1775
Coelolepida Nielsen & Kristensen, 1996
Myoglossata Kristensen & Nielsen, 1981
Neolepidoptera Packard, 1895
Infraorder Heteroneura Tillyard, 1918
Eulepidoptera Kiriakoff, 1948
Ditrysia Börner, 1925
Apoditrysia Minet, 1983, sedis mutabilis
Obtectomera Minet, 1986, sedis mutabilis
Macroheterocera Chapman, 1893, sedis mutabilis
Superfamily Noctuoidea Latreille, 1809, sedis mutabilis
"quadrifids"
Family Noctuidae Latreille, 1809 - owlet moths
Subfamily Acontiinae Guenée, 1841
Genus Abacena Walker, 1865
Genus Acaenica Hampson, 1918 [monotypic]
Genus Acidaliodes Hampson, 1910
Genus Acontarache Berio, 1977 [monotypic]
Genus Acremma Berio, 1959 [monotypic]
Genus Acropserotarache Berio, 1937 [monotypic]
Genus Aegleoides Berio, 1937 [monotypic]
Genus Alikangiana Strand, 1920 [monotypic]
Genus Alypophanes Turner, 1908
Genus Amphodia Möschler, 1880 [monotypic]
Genus Amynodes Warren, 1913
Genus Anateinoma Möschler, 1889 [monotypic]
[Genus Angitia Walker, 1858, nom. illeg.]
see Genus Blancharditia Biezanko & Ruffinelli, 1963
[Genus Anthophila Ochsenheimer, 1816, nom. illeg.]
see Genus Thalpochares Lederer, 1853
Genus Araeopterella Dyar, 1914 [monotypic]
Genus Arasada Moore, 1885
Genus Arenarba Berio, 1950
Genus Argyrargenta Berio, 1939 [monotypic]
Genus Argyropasta Hampson, 1910 [monotypic]
Genus Aroana Bethune-Baker, 1906
Genus Aulotarache Hampson, 1910
Genus Barastrotia Warren, 1913 [monotypic]
Genus Berresa Walker, 1859
Genus Blancharditia Biezanko & Ruffinelli, 1963
Genus Bostrodes Hampson, 1910
Genus Bryocodia Hampson, 1910
Genus Cadiorapa Nye, 1975
Genus Callipyris Meyrick, 1891
Genus Calloruza Hampson, 1918 [monotypic]
Genus Callostrotia Hampson, 1914 [monotypic]
Genus Calymma Hübner, 1823 [monotypic]
Genus Cardiosace Hampson, 1910
Genus Carmara Walker, 1863 [monotypic]
Genus Cautaeschra Hampson, 1910
Genus Cautatha Hampson, 1910 [monotypic]
Genus Ceratostrotia Warren, 1913 [monotypic]
Genus Cerynea Walker, 1859
Genus Chalenata Walker, 1864
[Genus Chamyris Guenée, 1852]
see Genus Cerma Hübner, 1818
Genus Chamyrisilla Draudt, 1950 [monotypic]
Genus Chara Staudinger, 1892
Genus Charoblemma Dyar, 1914
Genus Chelichares Hampson, 1910 [monotypic]
Genus Chionoxantha Hampson, 1914
Genus Chlorocodia Hampson, 1910 [monotypic]
Genus Chobata Walker, 1857 [monotypic]
Genus Cingalesa Hampson, 1894 [monotypic]
Genus Clytoscopa Turner, 1931
Genus Coccidiphaga Spuler, 1907 [monotypic]
Genus Codalithia Holloway, 1979
Genus Coeloturatia Strand, 1928 [monotypic]
Genus Conochares Smith, 1905
Genus Cophanta Walker, 1864
Genus Corgatha Walker, 1858
Genus Corgathalia Berio, 1966 [monotypic]
Genus Cretonia Walker, 1866
Genus Cryphioides Berio, 1964 [monotypic]
Genus Daedalina Möschler, 1881 [monotypic]
Genus Decticryptis Hampson, 1910 [monotypic]
Genus Diplothecta Turner, 1920 [monotypic]
Genus Dischalis Hampson, 1918 [monotypic]
Genus Ditrogoptera Hampson, 1898 [monotypic]
Genus Drobeta Walker, 1858
Genus Duhemia Rungs, 1943 [monotypic]
Genus Dymba Dyar, 1914 [monotypic]
Genus Dysgnathia Warren, 1913
Genus Ecnomia Turner, 1936 [monotypic]
Genus Ectrogatha Hampson, 1910 [monotypic]
Genus Enea Walker, 1864[1863] [monotypic]
Genus Enispades Bethune-Baker, 1911
Genus Enispodes Warren, 1913 [monotypic]
Genus Epicerynea Hampson, 1914 [monotypic]
Genus Epigrypera Hampson, 1910
Genus Epopsima Turner, 1920 [monotypic]
Genus Erastroides Hampson, 1893
Genus Eublarginea Berio, 1966 [monotypic]
Genus Eublemmistis Hampson, 1902
Genus Eublemmoides Bethune-Baker, 1906
Genus Eugatha Hampson, 1911 [monotypic]
Genus Eugnathia Warren, 1912
Genus Eugraptoblemma Warren, 1913
Genus Eulocastra Butler, 1886
Genus Eupsoropsis Berio, 1969 [monotypic]
Genus Eusimplex Berio, 1977
Genus Farara Bethune-Baker, 1908 [monotypic]
Genus Flammona Walker, 1864[1863]
Genus Formosamyna Strand, 1920 [monotypic]
Genus Fredina Brandt, 1939 [monotypic]
Genus Fruva Grote, 1877
Genus Fulvarba Berio, 1950 [monotypic]
Genus Gelenipsa Dyar, 1914 [monotypic]
Genus Gerarctia Hampson, 1905 [monotypic]
Genus Giubicolanta Berio, 1937 [monotypic]
Genus Glaucicodia Hampson, 1910 [monotypic]
Genus Goniophylla Turner, 1945 [monotypic]
Genus Gonophaea Hampson, 1910 [monotypic]
Genus Gyrohypsoma Staudinger, 1888 [monotypic]
Genus Habrophyes Turner, 1920 [monotypic]
Genus Haemaphlebia Hampson, 1910
Genus Haematosticta Hampson, 1896 [monotypic]
Genus Haplopseustis Meyrick, 1902 [monotypic]
Genus Haplostola Möschler, 1890 [monotypic]
Genus Heliocontia Hampson, 1910
Genus Hemispragueia Barnes & Benjamin, 1923 [monotypic]
Genus Heterorta Warren, 1913 [monotypic]
Genus Hiccoda Moore, 1882
Genus Himerois Turner, 1902
Genus Holocryptis Lucas, 1892
Genus Hoplotarache Hampson, 1910
Genus Hopothia Dyar, 1914 [monotypic]
Genus Hypangitia Hampson, 1918 [monotypic]
Genus Hypercodia Hampson, 1910
Genus Hypersophtha Berio, 1954
Genus Hypobleta Turner, 1908
Genus Hyposada Hampson, 1910
Genus Hyriodes Hampson, 1910 [monotypic]
Genus Ilattia Walker, 1858
Genus Lambana Walker, 1865
Genus Lamprolopha Hampson, 1914 [monotypic]
Genus Lepidopyrga Warren, 1914 [monotypic]
Genus Ligidia Walker, 1862 [monotypic]
Genus Lobocheilos Hampson, 1891, nom. illeg.
Genus Lochia Walker, 1865 [monotypic]
Genus Lophocerynea Hampson, 1918 [monotypic]
Genus Lophocryptis Hampson, 1914 [monotypic]
Genus Lophocyttarra Hampson, 1914
Genus Lophomilia Warren, 1913
Genus Lophorache Hampson, 1910
Genus Lophoruza Hampson, 1910
[Genus Lycaugesia Hampson, 1910, nom. illeg.]
see Genus Quandara Nye, 1975
Genus Lycaugesia Dognin, 1910
Genus Meizoglossa Hulstaert, 1924 [monotypic]
Genus Melaleucantha Draudt, 1950 [monotypic]
Genus Melamera Hampson, 1910 [monotypic]
Genus Meneptera Hampson, 1910 [monotypic]
Genus Mesembreosa Hulstaert, 1924 [monotypic]
Genus Mesocopsis Warren, 1913 [monotypic]
Genus Mesophractias Warren, 1913
Genus Mesoruza Warren, [1913] [monotypic]
[Genus Mestleta Walker, 1865]
Genus Metacausta Hampson, 1910
Genus Metachrostis Hübner, [1820]
Genus Metaemene Hampson, 1910
Genus Metapioplasta Wallengren, 1865
Genus Metasada Hampson, 1910
Genus Metaxyllia Dyar, 1922 [monotypic]
Genus Micardia Butler, 1878
[Genus Micra]
Genus Micracontia Hampson, 1895 [monotypic]
Genus Micraeschus Butler, 1878
Genus Micrantha Hampson, 1910
Genus Microedma Warren, 1913 [monotypic]
Genus Microphaea Hampson, 1910
Genus Microraphe Berio, 1937 [monotypic]
Genus Mictochroa Druce, 1909
[Genus Mictochroa Hampson, 1910, nom. illeg.]
see Genus Mictochroa Druce, 1909
Genus Mimasura Hampson, 1910
Genus Mimoruza Hampson, 1895 [monotypic]
Genus Monocymia Hampson, 1910 [monotypic]
Genus Nanamonodes Hampson, 1914
Genus Narangodes Hampson, 1910
Genus Neocerynea Hampson, 1918 [monotypic]
Genus Neochrostis Hampson, 1902 [monotypic]
Genus Neocodia Schaus, 1911
Genus Neoerastria McDunnough, 1937
Genus Nerastria McDunnough, 1937
Genus Niaccaba Walker, 1865 [monotypic]
Genus Obarza Berio, 1950 [monotypic]
Genus Oederastria Hampson, 1902 [monotypic]
Genus Oediblemma Hampson, 1918 [monotypic]
Genus Oedicodia Hampson, 1910
Genus Oenoptera Hampson, 1910
Genus Ortholeuca Hampson, 1910 [monotypic]
Genus Orthoruza Warren, 1913 [monotypic]
Genus Oruza Walker, 1862
Genus Oruzodes Warren, 1913
Genus Outaya Chrétien, 1911 [monotypic]
Genus Oxytrita Warren, 1913 [monotypic]
Genus Ozana Berio, 1950 [monotypic]
Genus Ozopteryx Saalmüller, 1891 [monotypic]
Genus Palura Walker, 1861
Genus Paracodia Druce, 1909
Genus Parangitia Druce, 1909
[Genus Parangitia Hampson, 1910, nom. illeg.]
see Genus Parangitia Druce, 1909
Genus Parasada Hampson, 1910
Genus Parerastria Warren, 1914 [monotypic]
Genus Paroruza Hampson, 1902
Genus Paurosceles Turner, 1945 [monotypic]
Genus Paventia Rossi, 1958 [monotypic]
Genus Pediarcha Turner, 1936 [monotypic]
Genus Penisa Warren, 1912
Genus Pennalticola Berio, 1973 [monotypic]
Genus Peperita Hampson, 1910 [monotypic]
Genus Perynea Hampson, 1910
Genus Phaeozona Hampson, 1910 [monotypic]
Genus Phyllophila Guenée, 1852
Genus Pluxilloides Berio, 1944 [monotypic]
Genus Prasinopyra Hampson, 1914
Genus Procriosis Hampson, 1910
Genus Prodosia Dyar, 1914 [monotypic]
Genus Prorachia Hampson, 1909 [monotypic]
Genus Proschora Turner, 1945 [monotypic]
Genus Protarache Hampson, 1910
Genus Protocryphia Barnes & McDunnough, 1918
Genus Pseudacidalia Hampson, 1894
Genus Pseudomicrodes Hampson, 1910
Genus Pseudophyllophila Berio, 1977 [monotypic]
Genus Pseudosiccia Roepke, 1956 [monotypic]
Genus Pseudozarba Warren, 1913
Genus Ptyopterota Hampson, 1894 [monotypic]
Genus Pyraloides Rebel, 1948 [monotypic]
Genus Pyripnoa Turner, 1920
Genus Rhodotarache Warren, 1914 [monotypic]
Genus Rhynchodia Hampson, 1910 [monotypic]
Genus Scotostena Hampson, 1910
Genus Silda Walker, 1863
Genus Smicroloba Warren, 1913
Genus Sophta Walker, 1862
Genus Steganiodes Hampson, 1910 [monotypic]
Genus Stenocodia Hampson, 1910 [monotypic]
Genus Stenocryptis Warren, 1913 [monotypic]
Genus Stenoloba Staudinger, 1892
Genus Stenorache Hampson, 1910 [monotypic]
Genus Stenostygia Hampson, 1910 [monotypic]
Genus Stylorache Hampson, 1910 [monotypic]
Genus Sugia Ueda, 1984
Genus Swinhoea Hampson, 1894 [monotypic]
Genus Syngatha Bethune-Baker, 1913
Genus Synthaca Turner, 1908 [monotypic]
Genus Tabomeeres Nye, 1975 [monotypic]
Genus Tarachidia Hampson, 1898
Genus Taraconica Berio, 1959
Genus Taseopteryx Butler, 1883
Genus Tavila Walker, 1869 [monotypic]
Genus Technemon Turner, 1945 [monotypic]
Genus Tegiapa Nye, 1975 [monotypic]
Genus Tephrochares Zerny, 1933 [monotypic]
Genus Tesomonoda Nye, 1975 [monotypic]
Genus Teucocranon Berio, 1937 [monotypic]
Genus Thalerastria Staudinger, 1897
[Genus Thalpochares Lederer, 1853]
Genus Thaumasiodes Turner, 1939 [monotypic]
Genus Therasea Grote, 1895
[Genus Thermesia Hübner, 1823]
Genus Thyatirina Hampson, 1910 [monotypic]
Genus Toanodes Warren, [1913] [monotypic]
Genus Toanopsis Prout, 1926 [monotypic]
Genus Trichogatha Warren, [1913]
[Genus Trichotarache Grote, 1875 [monotypic]]
see Genus Acontia Ochsenheimer, 1816
Genus Trissernis Meyrick, 1902
Genus Trogatha Hampson, 1910
Genus Trogoblemma Hampson, 1910
Genus Trogocraspis Hampson, 1918 [monotypic]
Genus Trogotorna Hampson, 1910 [monotypic]
Genus Uncula Swinhoe, 1900
Genus Uniptena Nye, 1975 [monotypic]
Genus Vescisa Walker, 1864
Genus Via Dyar, 1914 [monotypic]
Genus Xanthograpta Hampson, 1910
Genus Xanthomera Hampson, 1914 [monotypic]
[Genus Xanthoptera]
Genus Zurobata Walker, 1865
1 Tribe Acontiiniᵀ Guenée, 1841