Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Deuterostomia Grobben, 1908
Phylum Chordata Bateson, 1885 - chordates
Subphylum Vertebrata Cuvier, 1812 - vertebrates
Myopterygii - myopterygians
Superclass Gnathostomata Gegenbaur, 1874 - jawed vertebrates
Grade Teleostomi Bonaparte, 1836
Euteleostomi - bony vertebrates
Clade Sarcopterygii Romer, 1955
[crown] Rhipidistia Cope, 1887
Clade Tetrapodomorpha Ahlberg, 1991
Clade Osteolepidimorpha
Clade Tetrapoda Hatschek & Cori 1896 [Gauthier, Kluge & Rowe 1988]
[crown] Clade Neotetrapoda
Class Amphibia Gray, 1825
Infraclass Lissamphibia Haeckel, 1866
Batrachia Latreille, 1800
Order Anura Fischer von Waldheim, 1813
Lalagobatrachia D.R. Frost et al., 2006
Sokolanura D.R. Frost et al., 2006
Acosmanura Savage, 1973
Neobatrachia Reig, 1958
Phthanobatrachia D.R. Frost et al., 2006
Hyloides D.R. Frost et al., 2006
Notogaeanura Frost et al., 2006
Nobleobatrachia D.R. Frost et al., 2006
Meridianura D.R. Frost et al., 2006
Athesphatanura D.R. Frost et al., 2006
Leptodactyliformes D.R. Frost et al., 2006
Chthonobatrachia D.R. Frost et al., 2006
Hesticobatrachia D.R. Frost et al., 2006
Agastorophrynia D.R. Frost et al., 2006
Family Bufonidae Gray, 1825
Genus Atelopus Duméril & Bibron, 1841
[Atelopus leoperezii Ruiz-Carranza, Ardila-Robayo & Hernández-Camacho, 1994]
[Atelopus proboscideus (Boulenger, 1882)]
see Rhinella boulengeri Chaparro, Pramuk, Gluesenkamp & Frost, 2007
[Atelopus stelzneri (Weyenbergh, 1875)]
see Melanophryniscus stelzneri (Weyenbergh, 1875)
[Atelopus willimani Donoso-Barros, 1969]
01 Atelopus andinus Rivero, 1968
02 Atelopus angelito Ardila-Robayo & Ruiz-Carranza, 1998
03 Atelopus arsyecue Rueda-Almonacid, 1994
04 Atelopus arthuri Peters, 1973
05 Atelopus balios Peters, 1973
06 Atelopus bomolochos Peters, 1973
07 Atelopus boulengeri Peracca, 1904
08 Atelopus carauta Ruiz-Carranza & Hernández-Camacho, 1978
09 Atelopus carbonerensis Rivero, 1974
10 Atelopus carrikeri Ruthven, 1916
11 Atelopus certus Barbour, 1923
12 Atelopus chiriquiensis Shreve, 1936
13 Atelopus chirripoensis Savage & Bolaños, 2009
14 Atelopus chocoensis Lötters, 1992
15 Atelopus chrysocorallus La Marca, 1996
16 Atelopus coynei Miyata, 1980
17 Atelopus cruciger (Lichtenstein & Martens, 1856)
18 Atelopus dimorphus Lötters, 2003
19.1 Atelopus ebenoides Rivero, 1963
19.2 Atelopus marinkellei Cochran & Goin, 1970
20 Atelopus elegans (Boulenger, 1882)
21 Atelopus epikeisthos Lötters, Schulte & Duellman, 2005
22 Atelopus erythropus Boulenger, 1903
23 Atelopus eusebianus Rivero & Granados-Díaz, 1993
24 Atelopus eusebiodiazi Venegas, Catenazzi, Siu-Ting & Carrillo, 2008
25 Atelopus exiguus Boettger, 1892
26 Atelopus famelicus Rivero & Morales, 1995
27 Atelopus farci Lynch, 1993
28 Atelopus flavescens Duméril & Bibron, 1841
29 Atelopus franciscus Lescure, 1974
30 Atelopus galactogaster Rivero & Serna, 1993
31 Atelopus glyphus Dunn, 1931
32 Atelopus guanujo Coloma, 2002
33 Atelopus guitarraensis Osorno-Muñoz, Ardila-Robayo & Ruiz-Carranza, 2001
34 Atelopus halihelos Peters, 1973
35 Atelopus ignescens (Cornalia, 1849)
36 Atelopus laetissimus Ruiz-Carranza, Ardila-Robayo & Hernández-Camacho, 1994
37 Atelopus limosus Ibáñez, Jaramillo & Solís, 1995
38 Atelopus longibrachius Rivero, 1963
39 Atelopus longirostris Cope, 1868
40 Atelopus lozanoi Osorno-Muñoz, Ardila-Robayo & Ruiz-Carranza, 2001
41 Atelopus lynchi Cannatella, 1981
42 Atelopus mandingues Osorno-Muñoz, Ardila-Robayo & Ruiz-Carranza, 2001
43 Atelopus mindoensis Peters, 1973
44 Atelopus minutulus Ruiz-Carranza, Hernández-Camacho & Ardila-Robayo, 1988
45 Atelopus mittermeieri Acosta-Galvis, Rueda-Almonacid, Velásquez-Álvarez, Sánchez-Pacheco & Peña-Prieto, 2006
46 Atelopus monohernandezii Ardila-Robayo, Osorno-Muñoz & Ruiz-Carranza, 2002
47 Atelopus mucubajiensis Rivero, 1974
48 Atelopus muisca Rueda-Almonacid & Hoyos, 1992
49 Atelopus nahumae Ruiz-Carranza, Ardila-Robayo & Hernández-Camacho, 1994
50 Atelopus nanay Coloma, 2002
51 Atelopus nepiozomus Peters, 1973
52 Atelopus nicefori Rivero, 1963
53 Atelopus onorei Coloma, Lötters, Duellman & Miranda-Leiva, 2007
54 Atelopus oxapampae Lehr, Lötters & Lundberg, 2008
55 Atelopus oxyrhynchus Boulenger, 1903
56 Atelopus pachydermus (Schmidt, 1857)
57 Atelopus palmatus Andersson, 1945
58 Atelopus patazensis Venegas, Catenazzi, Siu-Ting & Carrillo, 2008
59 Atelopus pedimarmoratus Rivero, 1963
60 Atelopus peruensis Gray & Cannatella, 1985
61 Atelopus petersi Coloma, Lötters, Duellman & Miranda-Leiva, 2007
62 Atelopus petriruizi Ardila-Robayo, 1999
63 Atelopus pictiventris Kattan, 1986
64 Atelopus pinangoi Rivero, 1982
65 Atelopus planispina Jiménez de la Espada, 1875
66 Atelopus pulcher Boulenger, 1882
67 Atelopus pyrodactylus Venegas & Barrio, 2006
68 Atelopus quimbaya Ruiz-Carranza & Osorno-Muñoz, 1994
69 Atelopus reticulatus Lötters, Haas, Schick & Böhme, 2002
70 Atelopus sanjosei Rivero & Serna, 1989
71 Atelopus seminiferus Cope, 1874
72 Atelopus senex Taylor, 1952
73 Atelopus sernai Ruiz-Carranza & Osorno-Muñoz, 1994
74 Atelopus simulatus Ruiz-Carranza & Osorno-Muñoz, 1994
75 Atelopus siranus Lötters & Henzl, 2000
76 Atelopus sonsonensis Vélez-Rodriguez & Ruiz-Carranza, 1997
77 Atelopus sorianoi La Marca, 1983
78.1 Atelopus spumarius Cope, 1871
78.2 Atelopus barbotini Lescure, 1981
78.2.0 ["Atelopus babortini" nom. null.]
see Atelopus barbotini Lescure, 1981
78.3 Atelopus hoogmoedi Lescure, 1974
79 Atelopus spurrelli Boulenger, 1914
80 Atelopus subornatus Werner, 1899
81 Atelopus tamaense La Marca, García-Pérez & Renjifo, 1990
81.0 ["Atelopus tamaensis" La Marca et al., 1990]
see Atelopus tamaense La Marca, García-Pérez & Renjifo, 1990
82 Atelopus tricolor Boulenger, 1902
83 Atelopus varius (Lichtenstein & Martens, 1856)
83.0 ["Atelopus varius" Stannius, 1856, nom. nud.]
see Atelopus varius (Lichtenstein & Martens, 1856)
84 Atelopus vogli Müller, 1934
85 Atelopus walkeri Rivero, 1963
86 Atelopus zeteki Dunn, 1933 - Panamanian golden frog