Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Subphylum Chelicerata Heymons, 1901
Euchelicerata Weygoldt & Paulus, 1979
Prosomapoda Lamsdell, 2013
Planaterga Lamsdell, 2013
Dekatriata Lamsdell, 2013
Sclerophorata Kamenz, Staude & Dunlop, 2011
Class Arachnida Lamarck, 1801
Subclass Dromopoda Shultz, 1990 [paraphyletic]
Stomothecata Shultz, 2007, sedis mutabilis
Order Scorpiones Koch, 1837 - scorpions
Suborder Neoscorpionina Thorell & Lindström, 1885
Infraorder Orthosterni Pocock, 1911
Parvorder Buthida Soleglad & Fet, 2003
Superfamily Buthoideaᵀ Koch, 1837 [monotypic]
Family Buthidaeᵀ Koch, 1837
001 Genus †Archaeoananteroides Lourenço, 2016
002 Genus †Palaeoakentrobuthus Lourenço & Weitschat, 2000
003 Genus †Palaeoananteris Lourenço & Weitschat, 2001
004 Genus †Palaeogrosphus Lourenço, 2000
005 Genus †Palaeoisometrus Lourenço & Weitschat, 2005
006 Genus †Palaeolychas Lourenço & Weitschat, 1996
007 Genus †Palaeoprotobuthus Lourenço & Weitschat, 2000
008 Genus †Palaeospinobuthus Lourenço, Henderickx & Weitschat, 2005
009 Genus †Palaeotityobuthus Lourenço & Weitschat, 2000
010 Genus †Uintascorpio Perry, 1995
011 Genus Afroisometrus Kovařík, 1997
012 Genus Akentrobuthus Lamoral, 1976
013 Genus Alayotityus Armas, 1973
014 Genus Ananteris Thorell, 1891
015 Genus Ananteroides Borelli, 1911
016 Genus Androctonus Ehrenberg, 1828
017 Genus Anomalobuthus Kraepelin, 1900
018 Genus Apistobuthus Finnegan, 1932
019 Genus Australobuthus Locket, 1990
020 Genus Babycurus Karsch, 1886
021 Genus Baloorthochirus Kovařík, 1996
022 Genus Barbaracurus Kovařík, Lowe & Šťáhlavský, 2018
023 Genus Birulatus Vachon, 1974
024 Genus Buthacus Birula, 1908
025 Genus Butheoloides Hirst, 1925
026 Genus Butheolus Simon, 1883
027 Genus Buthiscus Birula, 1905
028 Genus Buthoscorpio Werner, 1936
029 Genus Buthusᵀ Leach, 1815
030 Genus Centruroides Marx, 1890
031 Genus Chaneke Francke, Teruel & Santibáñez-Lopez, 2014
032 Genus Charmus Karsch, 1879
033 Genus Cicileiurus Teruel, 2007
034 Genus Cicileus Vachon, 1948
035 Genus Compsobuthus Vachon, 1949
036 Genus Congobuthus Lourenço, 1999
037 Genus Darchenia Vachon, 1977
038 Genus Egyptobuthus Lourenço, 1999
039 Genus Femtobuthus Lowe, 2010
040 Genus Gint Kovařík, Lowe, Pliskova & Šťáhlavský, 2013
041 Genus Grosphus Simon, 1880
042 Genus Hemibuthus Pocock, 1900
043 Genus Hemilychas Hirst, 1911
044 Genus Heteroctenus Pocock, 1893
045 Genus Himalayotityobuthus Lourenço, 1997
046 Genus Hottentotta Birula, 1908
047 Genus Iranobuthus Kovařík, 1997
048 Genus Ischnotelson Esposito, Yamaguti, Souza, Pinto-da-Rocha & Prendini, 2017
049 Genus Isometroides Keyserling, 1885
050 Genus Isometrus Ehrenberg, 1828
051 Genus Jaguajir Esposito, Yamaguti, Souza, Pinto-da-Rocha & Prendini, 2017
052 Genus Karasbergia Hewitt, 1914
053 Genus Kraepelinia Vachon, 1974
054 Genus Lanzatus Kovařík, 2001
055 Genus Leiurus Ehrenberg, 1828
056 Genus Liobuthus Birula, 1898
057 Genus Lissothus Vachon, 1948
058 Genus Lychas Koch, 1845
059 Genus Lychasioides Vachon, 1974
060 Genus Mauritanobuthus Qi & Lourenço, 2007
061 Genus Mesobuthus Vachon, 1950
062 Genus Mesotityus González-Sponga, 1981
063 Genus Microananteris Lourenço, 2003
064 Genus Microbuthus Kraepelin, 1898
065 Genus Microcharmus Lourenço, 1995
066 Genus Microtityus Kjellesvig-Waering, 1966
067 Genus Neobuthus Hirst, 1911
068 Genus Neogrosphus Lourenço, 1995
069 Genus Neoprotobuthus Lourenço, 2000
070 Genus Odontobuthus Vachon, 1950
071 Genus Odonturus Karsch, 1879
072 Genus Orthochiroides Kovařík, 1998
073 Genus Orthochirus Karsch, 1892
074 Genus Pantobuthus Lourenço & Duhem, 2009
075 Genus Parabuthus Pocock, 1890
076 Genus Pectinibuthus Fet, 1984
077 Genus Physoctonus Mello-Leitão, 1934
078 Genus Picobuthus Lowe, 2010
079 Genus Plesiobuthus Pocock, 1900
080 Genus Polisius Fet, Capes & Sissom, 2001
081 Genus Pseudolissothus Lourenço, 2001
082 Genus Pseudolychas Kraepelin, 1911
083 Genus Pseudouroplectes Lourenço, 1995
084 Genus Razianus Farzanpay, 1987
085 Genus Reddyanus Vachon, 1972
086 Genus Rhopalurus Thorell, 1876
087 Genus Saharobuthus Lourenço & Duhem, 2009
088 Genus Sassanidotus Farzanpay, 1987
089 Genus Somalibuthus Kovařík, 1998
090 Genus Somalicharmus Kovařík, 1998
091 Genus Thaicharmus Kovařík, 1995
092 Genus Tityobuthus Pocock, 1893
093 Genus Tityopsis Armas, 1974
094 Genus Tityus Koch, 1836
095 Genus Troglorhopalurus Lourenço, Baptista & Giupponi, 2004
096 Genus Troglotityobuthus Lourenço, 2000
097 Genus Uroplectes Peters, 1861
098 Genus Vachoniolus Levy, Amitai & Shulov, 1973
099 Genus Vachonus Tikader & Bastawade, 1983
100 Genus Xenobuthus Lowe, 2018
101 Genus Zabius Thorell, 1893