Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Panorpida
Amphiesmenoptera
Order Lepidoptera Linnaeus, 1758 - butterflies, moths
Suborder Glossata Fabricius, 1775
Coelolepida Nielsen & Kristensen, 1996
Myoglossata Kristensen & Nielsen, 1981
Neolepidoptera Packard, 1895
Infraorder Heteroneura Tillyard, 1918
Eulepidoptera Kiriakoff, 1948
Ditrysia Börner, 1925
Apoditrysia Minet, 1983, sedis mutabilis
Obtectomera Minet, 1986, sedis mutabilis
Macroheterocera Chapman, 1893, sedis mutabilis
Superfamily Noctuoidea Latreille, 1809, sedis mutabilis
"quadrifids"
Family Erebidae Leach, 1815
Subfamily Arctiinae Leach, 1815
Tribe Lithosiini Billberg, 1820
Genus Abrochocis Dyar, 1914 [monotypic]
Genus Acanthofrontia Hampson, 1910
Genus Acco Bethune-Baker, 1904 [monotypic]
Genus Achroosia Hampson, 1900 [monotypic]
Genus Aclytophanes Turner, 1940 [monotypic]
Genus Adoxosia Hampson, 1900
Genus Adrepsa Moore, 1879 [monotypic]
Genus Aeolosia Hampson, 1900
Genus Aethosia Hampson, 1900 [monotypic]
Genus Agkonia Dognin, 1894
Genus Aglossosia Hampson, 1900
Genus Agrisius Walker, 1855
Genus Agylloides Strand, 1912
Genus Alepista de Toulgoët, 1976 [monotypic]
Genus Allochrista Roepke, 1946 [monotypic]
Genus Amalodeta Meyrick, 1889
Genus Amplicincia Field, 1950
Genus Anaemosia Hampson, 1918 [monotypic]
Genus Anaene Dyar, 1914
Genus Anaphleps Rothschild, 1916 [monotypic]
Genus Anaphosia Hampson, 1903
Genus Anaulosia Schaus, 1911 [monotypic]
Genus Antona Walker, 1854
Genus Apaidia Hampson, 1900
Genus Apistosia Hübner, 1818
Genus Apogurea Watson, 1980 [monotypic]
Genus Apothosia Hampson, 1918 [monotypic]
Genus Aptilosia Schaus, 1911 [monotypic]
Genus Archilema Birket-Smith, 1965
Genus Archithosia Birket-Smith, 1965
Genus Ardonea Walker, 1854
Genus Ardonissa Dognin, 1907 [monotypic]
Genus Aresia Barnes & McDunnough, 1913 [monotypic]
Genus Areva Walker, 1854
Genus Arhabdosia Dyar, 1905 [monotypic]
Genus Arrhythmica Turner, 1940 [monotypic]
Genus Ascaptesyle Dyar, 1906
Genus Asparus Watson, 1980 [monotypic]
Genus Astacosia Toulgoët, 1958
Genus Asura Walker, 1854
Genus Asurgylla Kiriakoff, 1958 [monotypic]
Genus Asuridia Hampson, 1900
Genus Asuridoides Daniel, 1951
Genus Asythosia Birket-Smith, 1965 [monotypic]
Genus Atelophleps Turner, 1940 [monotypic]
Genus Ateucheta Turner, 1940 [monotypic]
Genus Atolmis Hübner, [1819]
Genus Avela Walker, 1856 [monotypic]
Genus Balbura Walker, 1854
Genus Barsinella Butler, 1878
Genus Birgorima Watson, 1980 [monotypic]
Genus Bitecta Heylaerts, 1891
Genus Blabioides Hampson, 1900 [monotypic]
Genus Blavia Walker, 1862 [monotypic]
Genus Boadicea Tams, 1930
Genus Boenasa Walker, 1864
Genus Brachiosia Hampson, 1900 [monotypic]
Genus Bryantia Schaus, 1922 [monotypic]
Genus Brycea Walker, 1854
Genus Byrsia Walker, 1864
Genus Calamidia Butler, 1877
Genus Callisthenia Hampson, 1900
Genus Caprimimodes Rothschild, 1913 [monotypic]
Genus Carcinopodia Hampson, 1900
Genus Carilephia Schaus, 1924 [monotypic]
Genus Castronia Jones, 1912 [monotypic]
Genus Castulo Walker, 1854
Genus Caulocera Hampson, 1900
Genus Celamodes Hampson, 1914 [monotypic]
Genus Chamaita Walker, 1862
Genus Chionaema Herrich-Schäffer, [1856]
Genus Chionosia Hampson, 1900
Genus Chiretolpis Watson, 1980
Genus Chlorogenia Meyrick, 1889
Genus Chrysaeglia Butler, 1877 [monotypic]
Genus Chrysaegliodes Strand, 1912 [monotypic]
Genus Chrysallactis Hampson, 1900 [monotypic]
Genus Chrysasura Hampson, 1914
Genus Chrysochlorosia Hampson, 1900
Genus Chrysomesia Hampson, 1903 [monotypic]
Genus Chrysorabdia Butler, 1877
Genus Chrysoscota Hampson, 1900
Genus Chrysozana Hampson, 1900 [monotypic]
Genus Cincia Walker, 1854
Genus Clemendana Schaus, 1929 [monotypic]
Genus Cloesia Hampson, 1900
Genus Conilepia Hampson, 1900 [monotypic]
Genus Costarcha Hampson, 1891 [monotypic]
Genus Cragia Birket-Smith, 1965
Genus Cristulosia de Toulgoët, 1958
Genus Crocodeta Hampson, 1914 [monotypic]
Genus Crocosia Hampson, 1914 [monotypic]
Genus Ctenane Swinhoe, 1905
Genus Ctenosia Hampson, 1900
Genus Cybosia Hübner, 1819 [monotypic]
Genus Cyclomilta Hampson, 1900 [monotypic]
Genus Cyclosiella Hampson, 1900 [monotypic]
Genus Cyclosodes Hampson, 1901 [monotypic]
Genus Cyclosticta Hampson, 1900 [monotypic]
Genus Damias Boisduval, 1832
Genus Darantasia Walker, 1859
Genus Darantasiella Roepke, 1943 [monotypic]
Genus Darantoides Hampson, 1900 [monotypic]
Genus Deloplotela Hulstaert, 1924 [monotypic]
Genus Deua Walker, 1864 [monotypic]
Genus Diaconisia Hampson, 1914 [monotypic]
Genus Diadesmola Hulstaert, 1924 [monotypic]
Genus Diarhabdosia Hampson, 1900
Genus Dichrostoptera Hulstaert, 1924 [monotypic]
Genus Diduga Moore, 1887
Genus Didymonyx Watson, 1980 [monotypic]
Genus Dipaenae Walker, 1854
Genus Disaulota Hampson, 1900 [monotypic]
Genus Disoidemata Hampson, 1900
Genus Dixanaene Dyar, 1914 [monotypic]
Genus Dohertya Hampson, 1894 [monotypic]
Genus Dolichesia Schaus, 1911
Genus Dotha Walker, 1864 [monotypic]
Genus Endrosa Hübner, [1819]
Genus Epeiromulona Field, 1952
Genus Epitalara Dyar, 1905
Genus Eriomastyx Rothschild & Jordan, 1905
Genus Euclemensoides Strand, 1922 [monotypic]
Genus Euconosia Watson, 1980 [monotypic]
Genus Eucyclopera Hampson, 1895
Genus Eudoliche Möschler, 1878
Genus Eugoa Walker, 1857
Genus Eugonosia Hampson, 1900 [monotypic]
Genus Eugraphosia Hampson, 1918 [monotypic]
Genus Euproctosia Hampson, 1914 [monotypic]
Genus Eurosia Hampson, 1900
Genus Eurozonosia Hampson, 1914
Genus Eurylomia Felder, 1874
Genus Eurynora Hampson, 1914
Genus Eurypeplella Hedicke, 1923 [monotypic]
Genus Eutane Walker, 1854
Genus Eutelesia Hampson, 1900
Genus Euthyone Watson, 1980
Genus Euzeugapteryx Dyar, 1905 [monotypic]
Genus Exilisia de Toulgoët, 1958
Genus Fabresema Holloway, 1979
Genus Gampola Moore, 1878 [monotypic]
Genus Garudinia Moore, 1882
Genus Garudinistis Hampson, 1900
Genus Garudinodes Bethune-Baker, 1908
Genus Gaudeator Dyar, 1914 [monotypic]
Genus Geridixis Dyar, 1914 [monotypic]
Genus Geriojennsa Watson, 1980 [monotypic]
Genus Glaucosia Hampson, 1900 [monotypic]
Genus Graphosia Hampson, 1900
Genus Graptasura Hampson, 1900 [monotypic]
Genus Grucia Dyar, 1916 [monotypic]
Genus Gylla Bryk, 1953 [monotypic]
Genus Gymnasura Hampson, 1900
Genus Gymnochroma Hampson, 1900
Genus Habrochroma Turner, 1940 [monotypic]
Genus Halone Walker, 1854
Genus Halurgia Herrich-Schäffer, 1856 [monotypic]
Genus Hanoisiella De Joannis, 1928 [monotypic]
Genus Hassleria Jørgensen, 1934 [monotypic]
Genus Hectobrocha Meyrick, 1886
Genus Heliorabdia Hampson, 1911 [monotypic]
Genus Heliosia Hampson, 1900
Genus Hemipsilia Hampson, 1900 [monotypic]
Genus Hemonia Walker, 1863
Genus Hestiarcha Meyrick, 1886 [monotypic]
Genus Hesychopa Turner, 1940 [monotypic]
Genus Heterallactis Meyrick, 1886
Genus Heterotropa Turner, 1940 [monotypic]
Genus Hobapromea Watson, 1980 [monotypic]
Genus Holochrea Hampson, 1900 [monotypic]
Genus Holocraspedon Hampson, 1893
Genus Hypagoptera Hampson, 1900 [monotypic]
Genus Hypareva Hampson, 1900
Genus Hypasura Daniel, 1952 [monotypic]
Genus Hypermaepha Hampson, 1900
Genus Hyperthagylla Hampson, 1900 [monotypic]
Genus Hypeugoa Leech, 1899 [monotypic]
Genus Hyposhada Hampson, 1909 [monotypic]
Genus Hyposiccia Hampson, 1900
Genus Idopterum Hampson, 1894
Genus Illice Walker, 1859
Genus Ionthas Hampson, 1914 [monotypic]
Genus Isorropus Butler, 1880
Genus Katmeteugoa van Eecke, 1920 [monotypic]
Genus Lacydoides Daniel, 1955 [monotypic]
Genus Lambula Walker, 1866
Genus Lambulodes Hampson, 1914
Genus Lambulosia Hampson, 1914 [monotypic]
Genus Lamprosiella Collins, 1962
Genus Lamprostola Hampson, 1900
Genus Lepidilema Aurivillius, 1910 [monotypic]
Genus Lepista Wallengren, 1863
Genus Leptopepla Hampson, 1900 [monotypic]
Genus Leucorhodia Hampson, 1900 [monotypic]
Genus Lexis Wallengren, 1863
Genus Licnoptera Meyrick, 1889
Genus Lithoprocris Hampson, 1900
Genus Lithosarctia Daniel, 1954 [monotypic]
Genus Lobilema Aurivillius, 1910 [monotypic]
Genus Lobobasis Hampson, 1896 [monotypic]
Genus Lomuna Field, 1952 [monotypic]
Genus Lymantriopsis Hampson, 1900 [monotypic]
Genus Lysceia Walker, 1854 [monotypic]
Genus Macaduma Walker, 1866
Genus Macadumosia Hampson, 1914 [monotypic]
Genus Machairophora Hampson, 1893 [monotypic]
Genus Macrobrochis Herrich-Schäffer, [1856]
Genus Macroptila Dognin, 1894
Genus Macrosia Hampson, 1900
Genus Mahensia Fryer, 1912 [monotypic]
Genus Malesia van Eecke, 1920 [monotypic]
Genus Mantala Walker, 1862 [monotypic]
Genus Marsypophora Felder, 1874
Genus Megalobosia Hampson, 1914 [monotypic]
Genus Melaneama Butler, 1877
Genus Melastrota Hampson, 1905
Genus Meneclia Grünberg, 1910 [monotypic]
Genus Metagylla Hampson, 1907 [monotypic]
Genus Metallosia Hampson, 1900
Genus Metalobosia Hampson, 1900
Genus Metareva Hampson, 1900
Genus Meterythrosia Hampson, 1900 [monotypic]
Genus Meteugoa Hampson, 1900
Genus Meteura Hampson, 1900
Genus Metexilisia de Toulgoët, 1958 [monotypic]
Genus Micrilema Hampson, 1903 [monotypic]
Genus Microhyle Hampson, 1905
Genus Micrommia Felder, 1874 [monotypic]
Genus Microtane Hampson, 1901 [monotypic]
Genus Miltasura Roepke, 1946 [monotypic]
Genus Miltochrista Hübner, 1819
Genus Mimulosia de Toulgoët, 1958
Genus Mintopola Hampson, 1900
Genus Mithuna Moore, 1878
Genus Monosyntaxis Swinhoe, 1901
Genus Mulona Walker, 1866
Genus Muxta Birket-Smith, 1965 [monotypic]
Genus Nanna Birket-Smith, 1965, nom. illeg.
Genus Narosodes Moore, 1887
Genus Neagylla Hampson, 1900 [monotypic]
Genus Neardonaea Gaede, 1925 [monotypic]
Genus Neasura Hampson, 1900
Genus Neasuroides Matsumura, 1927
Genus Neoblavia Hampson, 1900 [monotypic]
Genus Neobrocha Meyrick, 1886
Genus Neoduma Hampson, 1918
Genus Neomulona Hampson, 1914 [monotypic]
Genus Neoscaptia Hampson, 1900
Genus Neosiccia van Eecke, 1927 [monotypic]
Genus Neotalara Hampson, 1914 [monotypic]
Genus Neothyone Hampson, 1914
Genus Neozana Hampson, 1914 [monotypic]
Genus Nephelomilta Hampson, 1900 [monotypic]
Genus Nephelosia Hampson, 1900 [monotypic]
Genus Nesiotica Turner, 1917 [monotypic]
Genus Nilgiricola Hampson, 1918 [monotypic]
Genus Nipponasura Inoue, 1965 [monotypic]
Genus Nishada Moore, 1878
Genus Nodozana Hampson, 1900
Genus Nolinophanes Hulstaert, 1924 [monotypic]
Genus Novosia de Toulgoët, 1958 [monotypic]
Genus Nudaria Haworth, 1809
Genus Nudina Staudinger, 1887 [monotypic]
Genus Nudosia Hampson, 1900 [monotypic]
Genus Nudur Dyar, 1914 [monotypic]
Genus Nyctochroa Felder, 1874 [monotypic]
Genus Nyctosia Hampson, 1900
Genus Ochrota Hopffer, 1862
Genus Odozana Walker, 1864
Genus Oedaleosia Hampson, 1900
Genus Oeonistis Hübner, [1819]
Genus Oeonosia Hampson, 1914 [monotypic]
Genus Omiosia Hampson, 1900 [monotypic]
Genus Onychipodia Hampson, 1900
Genus Onymapata Hulstaert, 1924 [monotypic]
Genus Opsaroa Hampson, 1905 [monotypic]
Genus Oreopola Turner, 1940 [monotypic]
Genus Ovenna Birket-Smith, 1965
Genus Ovipennis Hampson, 1900
Genus Oxacme Hampson, 1894
Genus Pachasura Dognin, 1923 [monotypic]
Genus Pachycerosia Hampson, 1900
Genus Padenia Moore, 1882
Genus Padenodes Hampson, 1914
Genus Paidia Hübner, 1819
Genus Palaeopsis Hampson, 1893
Genus Palaeosia Hampson, 1900
Genus Palaeosiccia Hampson, 1900 [monotypic]
Genus Palaeotype Hampson, 1918 [monotypic]
Genus Palaeozana Dyar, 1914 [monotypic]
Genus Parabitecta Draeseke, 1926 [monotypic]
Genus Paracincia Field, 1950
Genus Paradohertya Bethune-Baker, 1904 [monotypic]
Genus Paradoxosia Hampson, 1918 [monotypic]
Genus Paragylla Hampson, 1900
Genus Paralithosia Daniel, 1954
Genus Paramulona Hampson, 1900
Genus Paraona Moore, 1878
Genus Paraonagylla Berio, 1939 [monotypic]
Genus Parapalosia Dyar, 1905 [monotypic]
Genus Parascolia Watson, 1980 [monotypic]
Genus Parashada Hampson, 1905 [monotypic]
Genus Parasiccia Hampson, 1900
Genus Paratalara Dyar, 1905 [monotypic]
Genus Paratype Felder, 1874
Genus Parelictis Meyrick, 1886 [monotypic]
Genus Paremonia Hampson, 1900
Genus Parexilisia de Toulgoët, 1958
Genus Parvicincia Field, 1950 [monotypic]
Genus Pasteosia Hampson, 1900
Genus Paulianosia de Toulgoët, 1958 [monotypic]
Genus Paurophleps Hampson, 1900 [monotypic]
Genus Pelosia Hübner, [1819]
Genus Phacusosia Hampson, 1911
Genus Phaeophlebosia Hampson, 1900 [monotypic]
Genus Phaeosia Hampson, 1900
Genus Phaulosia Hampson, 1900 [monotypic]
Genus Phenacomorpha Turner, 1940 [monotypic]
Genus Philenora Rosenstock, 1885
Genus Phlogomera Hampson, 1914 [monotypic]
Genus Physetocneme Felder, 1861 [monotypic]
Genus Pitane Walker, 1854 [monotypic]
Genus Pliniola Strand, 1912 [monotypic]
Genus Plumareola van Eecke, 1920 [monotypic]
Genus Poliodule Hampson, 1900
Genus Poliosia Hampson, 1900
Genus Porphyrochrysa Turner, 1940 [monotypic]
Genus Prepiella Hampson, 1900
Genus Prinasura Hampson, 1903 [monotypic]
Genus Procridia Hampson, 1900 [monotypic]
Genus Procrimima Hampson, 1900
Genus Progona Berg, 1882
Genus Prolobosia Hampson, 1914 [monotypic]
Genus Pronola Hampson, 1900
Genus Prosiccia Hampson, 1914 [monotypic]
Genus Protolithosia Röber, 1925 [monotypic]
Genus Proxhyle de Toulgoët, 1959
Genus Pseudilema Bethune-Baker, 1904 [monotypic]
Genus Pseudlepista Hampson, 1910
Genus Pseudoblabes Zeller, 1853 [monotypic]
Genus Pseudomacroptila Fleming, 1951 [monotypic]
Genus Pseudophanes Turner, 1940 [monotypic]
Genus Pseudoscaptia Hampson, 1914 [monotypic]
Genus Pusiola Wallengren, 1863
Genus Pusiolania Strand, 1912 [monotypic]
Genus Ranghana Moore, 1878 [monotypic]
Genus Rhagophanes Zeller, 1853 [monotypic]
Genus Rhodographa Hampson, 1900 [monotypic]
Genus Saozana Dyar, 1914 [monotypic]
Genus Scaphidriotis Turner, 1899 [monotypic]
Genus Scaptesyle Walker, 1854
Genus Schalodeta Hampson, 1914 [monotypic]
Genus Schistophleps Hampson, 1891
Genus Scoliacma Meyrick, 1886
Genus Scoliosia Hampson, 1914 [monotypic]
Genus Serincia Dyar, 1914
Genus Seripha Walker, 1854
Genus Setina Schrank, 1802 [monotypic]
Genus Siccia Walker, 1854
Genus Sicciaemorpha van Eecke, 1920 [monotypic]
Genus Siculifer Hampson, 1896 [monotypic]
Genus Siopastea Schaus, 1922 [monotypic]
Genus Snellenopsis van Eecke, 1920 [monotypic]
Genus Sozusa Wallengren, 1863
Genus Spatulosia de Toulgoët, 1965
Genus Stenarcha Hampson, 1900 [monotypic]
Genus Stenaulis Hampson, 1900 [monotypic]
Genus Stenilema Hampson, 1909
Genus Stenopterosia de Joannis, 1928 [monotypic]
Genus Stenoscaptia Hampson, 1900
Genus Stenosia Hampson, 1900 [monotypic]
Genus Sterrhosia de Joannis, 1928 [monotypic]
Genus Stictosia Hampson, 1900 [monotypic]
Genus Stigmatophora Staudinger, 1881
Genus Striosia Hampson, 1914 [monotypic]
Genus Sylescaptia van Eecke, 1920
Genus Symmetrodes Meyrick, 1886
Genus Syntomimorpha Hampson, 1900 [monotypic]
Genus Talara Walker, 1866
Genus Tampea Snellen, 1897
Genus Teinomastyx Hampson, 1914
Genus Teratopora Meyrick, 1889
Genus Tesma Birket-Smith, 1965
Genus Teulisna Walker, 1862
Genus Thallarcha Meyrick, 1886
Genus Thermeola Hampson, 1900
Genus Thermograpta Hampson, 1914 [monotypic]
Genus Threnosia Hampson, 1900
Genus Thumatha Walker, 1866
Genus Tigrioides Butler, 1877
Genus Tineopsis Felder, 1861 [monotypic]
Genus Tmetoptera Felder, 1874 [monotypic]
Genus Tortricosia Hampson, 1900 [monotypic]
Genus Torycus Herrich-Schäffer, 1866 [monotypic]
Genus Tospitis Walker, 1863
Genus Trichareva Hampson, 1914 [monotypic]
Genus Trischalis Hampson, 1894
Genus Trissobrocha Turner, 1914 [monotypic]
Genus Tropacme Hampson, 1894 [monotypic]
Genus Tuina Butler, 1877
Genus Turlinia de Toulgoët, 1976 [monotypic]
Genus Tylanthes Meyrick, 1889
Genus Urozana Hampson, 1900
Genus Utriculofera Hampson, 1893
Genus Vianania Orfila, 1954
Genus Viettesia de Toulgoët, 1959
Genus Vulmara Schaus, 1924 [monotypic]
Genus Xanthetis Hampson, 1900 [monotypic]
Genus Xantholopha Hampson, 1900 [monotypic]
Genus Yelva Birket-Smith, 1965 [monotypic]
Genus Zadadrina Kiriakoff, 1954 [monotypic]
Genus Zobida Birket-Smith, 1965 [monotypic]
Genus Zygaenosia Hampson, 1900
1 Subtribe Acsalina Bendib & Minet, 1999
2 Subtribe Eudesmiina Bendib & Minet, 1999
3 Subtribe Cisthenina Bendib & Minet, 1999
4 Subtribe Lithosiinaᵀ Billberg, 1820