Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Panorpida
Amphiesmenoptera
Order Lepidoptera Linnaeus, 1758 - butterflies, moths
Suborder Glossata Fabricius, 1775
Coelolepida Nielsen & Kristensen, 1996
Myoglossata Kristensen & Nielsen, 1981
Neolepidoptera Packard, 1895
Infraorder Heteroneura Tillyard, 1918
Eulepidoptera Kiriakoff, 1948
Ditrysia Börner, 1925
Apoditrysia Minet, 1983, sedis mutabilis
Superfamily Tortricoidea Latreille, 1802, sedis mutabilis [monotypic]
Family Tortricidaeᵀ Latreille, 1802 - leafroller moths
Subfamily Tortricinaeᵀ Latreille, 1802
Genus Accra Razowski, 1964
Genus Acleris Hübner, [1825]
Genus Acroceuthes Meyrick, 1881
Genus Acroplectis Meyrick, 1927
Genus Adoxophyes Meyrick, 1881
Genus Aeolostoma Meyrick, 1910
Genus Aesiocopa Zeller, 1877
Genus Aethes Billberg, 1820
Genus Aleimma Hübner, [1825]
Genus Alytopistis Meyrick, 1920
Genus Amboyna Razowski, 1964
Genus Amorbia Clemens, 1860
Genus Amphicoecia Razowski, 1975
Genus Anacrusis Zeller, 1877
Genus Anchicremna Meyrick, 1926
Genus Ancyroclepsis Diakonoff, 1976
Genus Aneuxanthis Le Marchand, 1933
Genus Anisogona Meyrick, 1881
Genus Anisotenes Diakonoff, 1952
Genus Anopina Obraztsov, 1962
Genus Anthophrys Diakonoff, 1960
Genus Antigraptis Meyrick, 1930
Genus Antiphrastis Meyrick, 1930
Genus Apateta Turner, 1926
Genus Aphelia Hübner, 1825
Genus Aphrozestis Meyrick, 1931
Genus Aphthonocosma Diakonoff, 1953
Genus Apinoglossa Saalmüller, 1890
Genus Aplastoceros Diakonoff, 1953
Genus Apotoforma Busck, 1934
Genus Apura Turner, 1916
Genus Archactenis Diakonoff, 1960
Genus Archepandemis Mutuura, 1978
Genus Archicnephasia Razowski, 1983
Genus Archidemis Diakonoff, 1967
Genus Archigraptis Razowski, 1964
Genus Arctephora Diakonoff, 1953
Genus Ardeutica Meyrick, 1913
Genus Argyrotaenia Stephens, 1852
Genus Aristocosma Meyrick, 1881
Genus Arizelana Diakonoff, 1953
Genus Arotrophora Meyrick, 1881
Genus Ascerodes Meyrick, 1905
Genus Asterolepis Razowski, 1964
Genus Astrosa Diakonoff, 1951
Genus Atelodora Meyrick, 1881
Genus Atteria Walker, 1863
Tribe Atterini
Genus Authomaema Turner, 1916
Genus Avaria Kocak, 1981
Genus Axioprepes Turner, 1945
Genus Balioxena Meyrick, 1912
Genus Barnardiella Turner, 1925
Genus Barygnathella Diakonoff, 1956
Genus Bathypluta Diakonoff, 1950
Genus Battalia Kocak, 1981
Genus Beryllophantis Meyrick, 1938
Genus Borboniella Diakonoff, 1957
Genus Borneogena Diakonoff, 1941
Genus Brachiolia Razowski, 1964
Genus Brachyvalva Diakonoff, 1960
Genus Bradleyella Zimmerman, 1978, nom. illeg.
Genus Branchophantis Meyrick, 1938
Genus Brongersmia Diakonoff, 1972
Genus Cacoecimorpha Obraztsov, 1954
Genus Callibryastis Meyrick, 1912
Genus Camadeniana Strand, 1915
Genus Campotenes Diakonoff, 1960
Genus Capnoptycha Meyrick, 1920
Genus Capua Stephens, 1834
Genus Carphomigma Diakonoff, 1953
Genus Catamacta Meyrick, 1911
Genus Cenopis Zeller, 1875
Genus Cerace Walker, 1863
Tribe Ceracini
Genus Ceramea Diakonoff, 1952
Genus Chersomorpha Meyrick, 1926
Genus Chionothremma Diakonoff, 1952
Genus Chiraps Diakonoff & Razowski, 1971
Genus Choanograptis Meyrick, 1938
Genus Choristenes Diakonoff, 1954
Genus Choristoneura Lederer, 1859
Genus Chresmarcha Meyrick, 1910
Genus Chrysoxena Meyrick, 1912
Genus Clepsiphyes Diakonoff, 1976
Genus Clepsis Guenée, 1845
Genus Cleptacaca Diakonoff, 1953
Genus Cnephasia Curtis, 1826
Genus Cnephasitis Razowski, 1965
Tribe Cochylini
Genus Coeloptera Turner, 1945
Genus Coelostathma Clemens, 1860
Genus Copidostoma Diakonoff, 1954
Genus Cornicacoecia Obraztsov, 1954
Genus Cornusaccula Diakonoff, 1960
Genus Cornuticlava Diakonoff, 1960
Genus Cosmiophrys Diakonoff, 1960
Genus Ctenopseustis Meyrick, 1885
Genus Cudonigera Powell & Obraztsov, 1977
Genus Curvisaccula Dugdale, 1966
Genus Cuspidata Diakonoff, 1960
Genus Daemilus Yasuda, 1972
Genus Danilevskiana Kuznetzov, 1973
Genus Decodes Obraztsov & Powell, 1961
Genus Deltinea Pastrana, 1961
Genus Deltobathra Meyrick, 1923
Genus Dentisociaria Kuznetsov, 1970
Genus Diactenis Meyrick, 1907
Genus Diactora Diakonoff, 1960
Genus Diadelomorpha Diakonoff, 1944
Genus Dicellitis Meyrick, 1908
Genus Dichelia Guenée, 1845
Genus Dichelopa Lower, 1901
Genus Digitosa Diakonoff, 1960
Genus Diphtheropyga Diakonoff, 1952
Genus Diplocalyptis Diakonoff, 1976
Genus Dipterina Meyrick, 1882
Genus Doloploca Hübner, 1825
Genus Doridostoma Diakonoff, 1973
Genus Dorithia Powell, 1964
Genus Drachmobola Meyrick, 1907
Genus Eana Billberg, 1820
Genus Eboda Walker, 1866
Genus Ebodina Diakonoff, 1967
Genus Ecclitica Meyrick, 1923
Genus Ecnomiomorpha Obraztsov, 1959
Genus Egogepa Razowski, 1977
Genus Elaeodina Meyrick, 1926
Genus Electracma Meyrick, 1906
Genus Electraglaia Diakonoff, 1976
Genus Enarmonodes Kuznetsov, 1968
Genus Epagoge Hübner, [1825]
Genus Epalxiphora Meyrick, 1882
Genus Epichorista Meyrick, 1909
Genus Epichoristodes Diakonoff, 1960
Genus Epicnephasia Danilevskii, 1963
Genus Epiphyas Turner, 1927
Genus Epitrichosma Lower, 1908
Genus Epitymbia Meyrick, 1882
Tribe Epitymbini
Genus Eremas Turner, 1945
Genus Ericodesma Dugdale, 1971
Genus Eulia Hübner, 1825
Tribe Euliini
Genus Eurydoxa Filipjev, 1930
Genus Eurythecta Meyrick, 1883
Genus Exapate Hübner, [1825]
Genus Furcinula Diakonoff, 1960
Genus Gelophaula Meyrick, 1923
Genus Geogepa Razowski, 1977
Genus Gephyraspis Diakonoff, 1960
Genus Glyphidoptera Turner, 1916
Genus Gnorismoneura Issiki & Stringer, 1932
Genus Goniotorna Meyrick, 1933
Genus Gynnidomorpha Turner, 1916
Genus Harmologa Meyrick, 1882
Genus Herotyda Razowski, 1971
Genus Heterochorista Diakonoff, 1952
Genus Hiceteria Diakonoff, 1953
Genus Histura Razowski, 1981
Genus Homalernis Meyrick, 1908
Genus Homona Walker, 1863
Genus Homonoides Diakonoff, 1960
Genus Homonopsis Kuznetsov, 1964
Genus Hydaranthes Meyrick, 1928
Genus Hypsidracon Meyrick, 1934
Genus Idolatteria Walsingham, 1913
Genus Immariana Park & Byun, 1991
Genus Ioditis Meyrick, 1938
Genus Isochorista Meyrick, 1881
Genus Isodemis Diakonoff, 1952
Genus Isotenes Meyrick, 1938
Genus Kawabea Razowski, 1965
Genus Labidosa Diakonoff, 1960
Genus Laciniella Dugdale, 1966
Genus Lambertiodes Diakonoff, 1959
Genus Lamyrodes Meyrick, 1910
Genus Latibulocrinis Tuck, 1985
Genus Leontochroma Walsingham, 1900
Genus Leptochroptila Diakonoff, 1939
Genus Leurogyia Common, 1965
Genus Lobogenesis Razowski, 1990
Genus Lopharcha Diakonoff, 1941
Genus Lophoprora Meyrick, 1930
Genus Lozotaeniodes Obraztsov, 1954
Genus Lumaria Diakonoff, 1976
Genus Macrothyma Diakonoff, 1952
Genus Mantua Zimmerman, 1978
Genus Maoritenes Dugdale, 1966
Genus Matronula Lepeletier & Serville, 1828
Genus Megalomacha Diakonoff, 1960
Genus Meridemis Diakonoff, 1976
Genus Meritastis Meyrick, 1910
Genus Merophyas Common, 1964
Genus Mesocalyptis Diakonoff, 1953
Genus Metachorista Meyrick, 1938
Genus Metamesia Diakonoff, 1960
Genus Metrernis Meyrick, 1906
Genus Mictoneura Meyrick, 1881
Genus Midaellobes Viette, 1990
Genus Mimeoclysia Diakonoff, 1941
Genus Neocalyptis Diakonoff, 1941
Genus Neosphaleroptera Réal, 1953
Genus Nesochoris Clarke, 1965
Genus Nesoscopa Meyrick, 1926
Genus Niasoma Busck, 1940
Genus Niphothixa Diakonoff, 1960
Genus Notioclepsis Diakonoff, 1983
Genus Ochetarcha Meyrick, 1924
Genus Oligotenes Diakonoff, 1954
Genus Orthocomotis Dognin, 1905
Genus Oxypteron Staudinger, 1871
Genus Palaeotoma Meyrick, 1881
Genus Panaphelix Walsingham, 1907
Genus Pandemis Hübner, 1825
Genus Pandurista Meyrick, 1918
Genus Panegyra Diakonoff, 1960
Genus Paracomotis Razowski, 1982
Genus Paracroesia Yasuda, 1972
Genus Paramesiodes Diakonoff, 1960
Genus Paranepsia Turner, 1916
Genus Paraphyas Turner, 1927
Genus Paraptila Meyrick, 1912
Genus Parastranga Meyrick, 1910
Genus Paratorna Meyrick, 1907
Genus Pareboda Razowski, 1966
Genus Pentacitrotus Butler, 1881
Genus Peraglyphis Common, 1963
Genus Periclepsis Bradley, 1977
Genus Peteliacma Meyrick, 1912
Genus Phaenacropista Diakonoff, 1941
Genus Phalonidia Le Marchand, 1935
Genus Philedone Hübner, [1825]
Genus Philedonides Obraztsov, 1954
Genus Philocryptica Meyrick, 1923
Genus Phlebozemia Diakonoff, 1984
Tribe Phricanthini
Genus Pisinidea Butler, 1883
Genus Planostocha Meyrick, 1912
Genus Planotortrix Dugdale, 1966
Genus Platynota Clemens, 1860
Genus Platysemaphora Diakonoff, 1960
Genus Polemograptis Meyrick, 1910
Genus Polydrachma Meyrick, 1928
Genus Proactenis Diakonoff, 1941
Genus Procalyptis Meyrick, 1910
Genus Procrica Diakonoff, 1960
Genus Proeulia Clarke, 1962
Genus Propiromorpha Obraztsov, 1955
Genus Proselena Meyrick, 1881
Genus Protarchella Diakonoff, 1956
Genus Prothelymna Meyrick, 1882
Genus Protopterna Meyrick, 1908
Genus Pseudargyrotoza Obraztsov, 1954
Genus Pseudeboda Razowski, 1964
Genus Pseudeulia Obraztsov, 1954
Genus Pseudocroesia Razowski, 1966
Genus Pseudomeritastis Obraztsov, 1966
Genus Pteridoporthis Meyrick, 1937
Genus Pternozyga Meyrick, 1908
Genus Ptycholomoides Obraztsov, 1954
Genus Punctapinella Brown, 1991
Genus Pyrgotis Meyrick, 1881
Genus Pyrsarcha Meyrick, 1932
Genus Rhabdotenes Diakonoff, 1960
Genus Rhomboceros Meyrick, 1910
Genus Rhopalotenes Diakonoff, 1960
Genus Rhynchophyllis Meyrick, 1932
Genus Rhythmologa Meyrick, 1926
Genus Sanguinograptis Razowski, 1981
Genus Schoenotenes Meyrick, 1908
Tribe Schoenotenini
Genus Sclerodisca Razowski, 1964
Genus Scolioplecta Meyrick, 1881
Genus Scotiophyes Diakonoff, 1976
Genus Scyphoceros Turner, 1925
Genus Scytalognatha Diakonoff, 1956
Genus Selenodes Guenée, 1845
Genus Sisurcana Powell, 1986
Genus Snodgrassia Diakonoff, 1967
Genus Sociosa Diakonoff, 1959
Genus Sorensenata Salmon & Bradley, 1956
Genus Sparganopseustis Powell & Lambert, 1986
Genus Sparganothis Hübner, 1825
Genus Sparganothoides Lambert & Powell, 1986
Genus Spatalistis Meyrick, 1907
Genus Sphaleroptera Guenée, 1845
Genus Spheterista Meyrick, 1912
Genus Stenarchella Diakonoff, 1967
Genus Stenotenes Diakonoff, 1954
Genus Subargyrotaenia Obraztsov, 1961
Genus Syllomatia Common, 1963
Genus Symphygas Common, 1963
Genus Syncratus Common, 1965
Genus Syndemis Hübner, 1825
Genus Synnoma Walsingham, 1879
Genus Synochoneura Obraztsov, 1955
Genus Taeniarchis Meyrick, 1931
Genus Tanychaeta Common, 1963
Genus Templemania Busck, 1931
Genus Terricula Fal'kovich, 1965
Genus Terthreutis Meyrick, 1918
Genus Thrincophora Meyrick, 1881
Genus Tinacrucis Powell, 1986
Genus Tracholena Common, 1965
Genus Transita Diakonoff, 1976
Genus Transtillaspis Razowski, 1987
Genus Tremophora Diakonoff, 1953
Genus Trophocosta Razowski, 1964
Genus Trychnophylla Turner, 1926
Genus Tymbarcha Meyrick, 1908
Genus Uelia Razowski, 1982
Genus Ulodemis Meyrick, 1907
Genus Vellonifer Razowski, 1964
Genus Vialonga Diakonoff, 1960
Genus Viettea Diakonoff, 1960
Genus Williella Horak, 1984
Genus Xeneda Diakonoff, 1961
Genus Xenophylla Diakonoff, 1960
Genus Xenotemna Powell, 1964
Genus Xenotenes Diakonoff, 1954
Genus Xenothictis Meyrick, 1910
Genus Xerocnephasia Leraut, 1979
Genus Zacorisca Meyrick, 1910
1 Tribe Tortriciniᵀ
2 Tribe Cnephasiini
3 Tribe Archipini
4 Tribe Sparganothini