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Reference: A.W.A. Kellner, 2003b. In E. Buffetaut & J.-M. Mazin (eds.), Geol. Soc. Lond. Spec. Publ. (no. 217)
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Article

Kellner, A.W.A. 2003b. Pterosaur phylogeny and comments on the evolutionary history of the group. In E. Buffetaut & J.-M. Mazin (eds.), Geol. Soc. Lond. Spec. Publ. (no. 217), 13 Oct 2003: [105]-137. ≡

New: Original clade name & spelling

†Asiaticognathidae (Clade) Kellner 2003Page: 111 ("n. taxon").
†Novialoidea (Clade) Kellner 2003 nom. cl. inval.Page: 115 ("n. taxon").

New: Subsequent clade name (orig. spell.)

†Anhangueridae (Clade) Campos & Kellner 1985 [Kellner 2003] nom. cl. inval.Page: 123 ("Campos & Kellner 1985").
†Anurognathidae (Clade) Kuhn 1937 [Kellner 2003] nom. cl. inval.Page: 108 ("Kuhn 1937").
†Azhdarchidae (Clade) Padian 1986 [Kellner 2003] nom. cl. inval.Page: 125 ("Nessov 1984").
†Ctenochasmatidae (Clade) Kuhn 1960 [Kellner 2003] nom. cl. inval.Page: 120 ("Nopcsa 1928").
†Dsungaripteridae (Clade) Young 1964 [Kellner 2003] nom. cl. inval.Page: 124 ("Young 1964").
†Gallodactylidae (Clade) Wellnhofer 1991 [Kellner 2003]Page: 120 ("Fabre 1974").
†Novialoidea (Clade) Kellner 2003 [Andres 2021]Page: tbl. 1 (p. 112: "Kellner (2003)").
†Nyctosauridae (Clade) Hay 1902 [Kellner 2003] nom. cl. inval.Page: 121 ("(=Nyctosaurinae) Williston 1903").
†Pterodactyloidea (Clade) Plieninger 1901 [Kellner 2003], nom. cl. inval.Page: 117.
†Rhamphorhynchidae (Clade) Hoernes 1884 [Kellner 2003] nom. cl. inval.Page: 117 ("Nopcsa 1928").
†Tapejaridae (Clade) Kellner 1989 [Kellner 2003] nom. cl. inval.Page: 125 ("Kellner 1989").

New: Original definition [PhyloCode]

†Anhangueridae (Clade) Campos & Kellner 1985 [Kellner 2003] nom. cl. inval.Page: 123 ("Definition.").
†Anurognathidae (Clade) Kuhn 1937 [Kellner 2003] nom. cl. inval.Page: 108 ("Definition.").
†Archaeopterodactyloidea (Clade) Kellner 1996 [Kellner 2003] nom. cl. inval.Page: 118 ("Kellner 1996", "Definition.").
†Asiaticognathidae (Clade) Kellner 2003Page: 111 ("Definition.").
†Azhdarchidae (Clade) Padian 1986 [Kellner 2003] nom. cl. inval.Page: 125 ("Definition.").
†Azhdarchoidea (Clade) Unwin 1995 [Kellner 2003] nom. cl. inval.Page: 125 ("Unwin 1995", "Definition.").
†Campylognathoididae (Clade) Wild 1984 [Kellner 2003]Page: 115 ("Kuhn 1967", "Definition.").
†Ctenochasmatidae (Clade) Kuhn 1960 [Kellner 2003] nom. cl. inval.Page: 120 ("Definition.").
†Dsungaripteridae (Clade) Young 1964 [Kellner 2003] nom. cl. inval.Page: 124 ("Definition.").
†Dsungaripteroidea (Clade) Young 1964 [Kellner 2003]Page: 120 ("Young 1964", "Definition.").
†Gallodactylidae (Clade) Wellnhofer 1991 [Kellner 2003]Page: 120 ("Definition.").
†Novialoidea (Clade) Kellner 2003 nom. cl. inval.Page: 115 ("Definition.").
†Nyctosauridae (Clade) Hay 1902 [Kellner 2003] nom. cl. inval.Page: 121 ("Definition.").
†Pteranodontoidea (Clade) Kellner 1996 [Kellner 2003] nom. cl. inval.Page: 122 ("Kellner 1996", "Definition.").
†Pterodactyloidea (Clade) Plieninger 1901 [Kellner 2003], nom. cl. inval.Page: 117 ("Definition.").
†Rhamphorhynchidae (Clade) Hoernes 1884 [Kellner 2003] nom. cl. inval.Page: 117 ("Definition.").
†Tapejaridae (Clade) Kellner 1989 [Kellner 2003] nom. cl. inval.Page: 125 ("Definition.").
†Tapejaroidea (Clade) Kellner 1996 [Kellner 2003]Page: 124 ("Kellner 1996", "Definition.").

New: Emended definition [PhyloCode]

†Ornithocheiroidea (Clade) Seeley 1876 [Bennett 1994] nom. cl. inval.Page: 122 ("Bennett 1994", "Definition."). [node-based].
†Pterosauria (Clade) Owen 1842 [Sereno 1991], nom. cl. inval.Page: 107 ("Kaup 1834", "Definition."). [node-based].

New: Description/Diagnosis in English

†Anhangueridae (Clade) Campos & Kellner 1985 [Kellner 2003] nom. cl. inval.Page: 123 ("Synapomorphies.").
†Azhdarchidae (Clade) Padian 1986 [Kellner 2003] nom. cl. inval.Page: 126 ("Synapomorphies.").
†Azhdarchoidea (Clade) Unwin 1995 [Kellner 2003] nom. cl. inval.Page: 125 ("Synapomorphies.").
†Campylognathoididae (Clade) Wild 1984 [Kellner 2003]Page: 115-116 ("Synapomorphies.").
†Ctenochasmatidae (Clade) Kuhn 1960 [Kellner 2003] nom. cl. inval.Page: 120 ("Synapomorphies.").
†Dsungaripteridae (Clade) Young 1964 [Kellner 2003] nom. cl. inval.Page: 124-125 ("Synapomorphies.").
†Dsungaripteroidea (Clade) Young 1964 [Kellner 2003]Page: 121 ("Synapomorphies.").
†Gallodactylidae (Clade) Wellnhofer 1991 [Kellner 2003]Page: 120 ("Synapomorphies.").
†Nyctosauridae (Clade) Hay 1902 [Kellner 2003] nom. cl. inval.Page: 121 ("Synapomorphies.").
†Pterodactyloidea (Clade) Plieninger 1901 [Kellner 2003], nom. cl. inval.Page: 117-118 ("Synapomorphies.").
†Rhamphorhynchidae (Clade) Hoernes 1884 [Kellner 2003] nom. cl. inval.Page: 117 ("Synapomorphies").
†Tapejaridae (Clade) Kellner 1989 [Kellner 2003] nom. cl. inval.Page: 125 ("Synapomorphies.").

Original Classification (root)

(Clade) †Pterosauria Owen 1842 [Sereno 1991], nom. cl. inval.

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Last updated: 28 Aug 2026