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Reference: C.E. ZoBell & H.C. Upham, 1944. Bull. Scripps Inst. Oceanogr. Univ. Calif. Tech. Ser. 5
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Zogg, H. & Upton, S.J. 1944. A List of Marine Bacteria Including Descriptions of Sixty New Species. Bull. Scripps Inst. Oceanogr. Univ. Calif. Tech. Ser. 5: 239-292.

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