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Reference: G.M. Garrity et al., 2007
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Garrity, G.M., Lilburn, T.G., Cole, J.R., Harrison, S.H., Euzéby, J. & Tindall, B.J. 2007. Part 10 - The Bacteria: Phylum "Actinobacteria": Class (Actinobacteria. Taxonomic Outline of the Bacteria and Archaea, Release 7.7., 6 Mar 2007: 399-539.

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