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Reference: G.M. Garrity et al., 2007
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Garrity, G.M., Lilburn, T.G., Cole, J.R., Harrison, S.H., Euzéby, J. & Tindall, B.J. 2007. Part 4 - The Bacteria: Phylum "Proteobacteria", Class Betaproteobacteria. Taxonomic Outline of the Bacteria and Archaea, Release 7.7., 6 Mar 2007: 112-147.

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