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Reference: C. Peña et al., 2006. Mol. Phyl. Evol. 40
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Peña, C., Wahlberg, N., Weingartner, E., Kodandaramaiah, U., Nylin, S., Freitas, A.V.L. & Brower, A.V.Z. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Mol. Phyl. Evol. 40: 29-49. [Online: 24 Mar 2006]

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