Whitehead, P.J.P., Nelson, G.J. & Wongratana, T. 1988. Part 2 - Engraulididae. Vol. 7. Clupeoid Fishes of the World (Suborder Clupeoidei). An Annotated and Illustrated Catalogue of the Herrings, Sardines, Pilchards, Sprats, Shads, Anchovies and Wolf-herrings. FAO Species Catalogue. FAO Fish. Synop. (125): 305-579. ≡