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Reference: D.M. Bramble & J.H. Hutchison, 2014
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Bramble, D.M. & Hutchison, J.H. 2014. Morphology, Taxonomy, and Distribution of North American Tortoises: An Evolutionary Perspective. In D.C. Rostal, E.D. McCoy & H.R. Mushinsky (eds.), Biology and Conservation of North American Tortoises. Johns Hopkins University Press, Baltimore: [1]-12.

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