Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade natates Cavalier-Smith, 2021
Clade photaria Cavalier-Smith, 2021
Clade eucorta Cavalier-Smith, 2021
Superkingdom Corticata Lankester, 1878 - corticates
Kingdom Plantae Haeckel (1866) - plants
Subkingdom Viridiplantae Caval.-Sm. (1981) - green plants
Infrakingdom Streptophyta Caval.-Sm. (1993)
[crown] Clade Phragmoplastophyta Lecointre & Le Guyader 2006 [Hall, McCourt, Delwiche, Mishler & Karol 2020]
[crown] Clade Embryophytaᵀ Engler 1887 [Mishler & Qiu 2020]
Clade Pan-Tracheophytaᵀ Cantino & Donoghue 2007 [Cantino & Donoghue 2020]
Clade Apo-Tracheophytaᵀ Cantino & Donoghue 2007 [Cantino & Donoghue 2020]
[crown] Phylum Tracheophytaᵀ Sinnott ex Caval.-Sm. (1998) - vascular plants
Clade Pan-Euphyllophytaᵀ Cantino & Donoghue 2007 [Cantino & Donoghue 2020]
[crown] Clade Euphyllophytaᵀ Lecointre & Le Guyader 2006 [Cantino & Donoghue 2020]
Clade Pan-Spermatophytaᵀ Cantino & Donoghue 2007 [Cantino, Doyle & Donoghue 2020]
Clade Lignophytaᵀ Crane & Kenrick 1997, nom. cl. inval.
Clade Apo-Spermatophytaᵀ Cantino & Donoghue 2007 [Cantino, Doyle & Donoghue 2020]
[crown] Subphylum Spermatophytinaᵀ Caval.-Sm. (1998) - seed plants
Clade Pan-Angiospermae Cantino & Donoghue 2007 [Doyle, Cantino & Donoghue 2020]
[crown] Class Magnoliopsidaᵀ Brongn. (1843)
[crown] Clade Mesangiospermaeᵀ Donoghue, Doyle & Cantino 2007 [Donoghue, Doyle & Cantino 2020]
Clade Tricolpataeᵀ Donoghue, Doyle & Cantino 2007 [Donoghue, Doyle & Cantino 2020]
[crown] Clade Eudicotyledoneaeᵀ Donoghue, Doyle & Cantino 2007 [Donoghue, Doyle & Cantino 2020]
[crown] Clade Gunneridaeᵀ Soltis, Soltis & Judd 2007 [Soltis, Soltis & Judd 2020]
[crown] Clade Pentapetalaeᵀ Soltis, Soltis & Judd 2007 [Soltis, Soltis & Judd 2020]
[crown] Clade Superasteridaeᵀ Judd, Soltis & Soltis 2011 [Judd, Soltis & Soltis 2020]
Superorder Asteranaeᵀ Takht. (1967)
[crown] Clade Gentianidaeᵀ Olmstead, Judd & Cantino 2007 [Olmstead, Judd & Cantino 2020]
[crown] Clade Lamiidaeᵀ Takhtajan 1987 [Olmstead, Judd & Cantino 2020]
[crown] Clade Lamianaeᵀ Takhtajan 1967 [Olmstead & Judd 2020]
Order Boraginalesᵀ Juss. ex Bercht. & J.Presl (1820), sedis mutabilis [monotypic]
Family Boraginaceaeᵀ Juss. (1789), nom. cons. - borage family
Genus Actinocarya Benth. (1876)
Genus Adelocaryum Brand
Genus Aegonychon Gray
Genus Afrotysonia S. Rauschert (1982)
Genus Amblynotus (A. de Candolle) Johnst. (1924)
Genus Amphibologyne Brand
Genus Ancistrocarya Maxim. (1872)
Genus Anoplocaryum Ledebour (1847)
Genus Antiotrema Handel-Mazzetti (1920)
Genus Antiphytum DC. ex C.F. Meisner (1840)
Genus Austrocynoglossum Popov ex R.R.Mill
Genus Benthamia Lindl.
Genus †Biorbia M.K. Elias (1932)
Genus †Borraginites Heer (1859)
Genus Bothriospermum Bunge (1833)
Genus Brachybotrys Maxim. ex Oliv. (1878)
Genus Brandella R.R. Mill (1986)
Genus †Calvarinus C. Reid & E.M. Reid (1910)
Genus Cerinthodes
Genus Chionocharis I.M.Johnst. (1924)
Genus Choriantha Riedl (1961)
Genus Craniospermum Lehm. (1818)
Genus Crucicaryum Brand (1929)
Genus Cynoglossopsis Brand, in Engl. (1931)
Genus Cynoglottis (Guşuleac) M. Vural & Kit Tan (1983)
Genus Cystostemon I.B. Balfour (1883)
Genus Dasynotus Johnst. (1948)
Genus †Davisella E.M. Reid & M.E.J. Chandler (1933)
Genus Echinospermum Sweet ex Lehm.
Genus Echiochilon Desf. (1798)
Genus †Ehretiaecarpum P. Menzel (1914)
Genus Embadium J.M.Black (1931)
Genus Gastrocotyle Bunge (1849)
Genus Gerascanthus P.Browne
Genus Glandora D. C. Thomas, Weigend & Hilger
Genus Gyrocaryum B. Valdes (1983)
Genus Harpagonella A.Gray (1876)
Genus Heterocaryum A.DC.
Genus Hilsenbergia Tausch ex Rchb.
Genus Kuschakewiczia Regel & Smirn.
Genus Lasiarrhenum I.M.Johnst. (1924)
Genus Lasiocaryum I.M.Johnst. (1925)
Genus Lepechiniella Popov (1953)
Genus Lindelofia Lehm. (1850)
Genus Lithocardium
Genus †Lithospermites E.W.Berry (1929)
Genus Lycopsis L. (1753)
Genus Macromeria D.Don (1832)
Genus Maharanga A.DC. (1846)
Genus Margarospermum (Rchb.) Opiz
Genus Metaeritrichium W.T.Wang
Genus Microcaryum I.M.Johnst. (1924)
Genus Microparacaryum (Popov ex Riedl) Hilger & Podlech
Genus Microula Bentham, in Bentham & Hook.f. (1876)
Genus Miltitzia A.DC. (1845)
Genus Mimophytum Greenman (1905)
Genus Morelosia Lex.
Genus Moritzia DC. ex C.F. Meisner (1840)
Genus †Myoporiphyllum Ettingsh. (1891)
Genus Myosotidium Hook. (1859)
Genus Myriopus Small
Genus Neatostema Johnst. (1953)
Genus Nesocaryum Johnst. (1927)
Genus Nomosa Johnst. (1954)
Genus Nyctelea Scop.
Genus Ogastemma R.K. Brummitt (1982)
Genus Omphalopus Naudin (1851)
Genus Omphalotrigonotis W.T.Wang
Genus Onosmodium Michx. (1803)
Genus Oreocarya Greene
Genus Pardoglossum Barbier & Mathez (1973)
Genus Patagonica Boehm.
Genus Pectocarya DC. ex Meisner (1840)
Genus Pentalophus A.DC.
Genus Pioctonon Raf.
Genus Pontechium Böhle & Hilger
Genus †Prolithospermum Elias (1942)
Genus Pseudomertensia H. Riedl, in K.H. Rechinger (1967)
Genus Psilolaemus I.M.Johnst. (1954)
Genus Schmidelia L.
Genus Setulocarya R.R.Mill & D.G.Long
Genus Sinojohnstonia H.H. Hu (1936)
Genus Stenosolenium Turcz.
Genus Thaumatocaryon Baill. (1890)
Genus Thyrocarpus Hance (1862)
Genus Varronia P.Browne
Genus Vaupelia Brand
Genus Viticella Dill. ex Moench
1 Genus Hoplestigma Pierre (1899)
2 Subfamily Cordioideae A.Gray
3 Subfamily Ehretioideae Arn.
4 Subfamily Heliotropioideae
5 Subfamily Hydrophylloideae Burnett
6 Subfamily Boraginoideaeᵀ (Juss.) Arn. (1832)
7 Subfamily Lennooideae