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Scarabaeinae (Subfamily) ..Ateuchini (Tribe) ..Canthonini (Tribe) ..Eucraniini (Tribe) ..Eurysternini (Tribe) ..Gymnopleurini (Tribe) ..Oniticellini (Tribe) ..Onitini (Tribe) ..Onthophagini (Tribe) ..Phanaeini (Tribe) ..Scarabaeini (Tribe) ..Sisyphini (Tribe) ..Coprini (Tribe) ....Copris (Genus) ....Coptodactyla (Genus) ....Oxyomus (Genus)

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Last updated: 1 Feb 2024