Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Domain Bacteria Woese et al. 2024
             Kingdom Pseudomonadati (Gibbons and Murray 1978) Oren and Göker 2024
               Phylum Thermodesulfobacteriota Garrity and Holt 2021
                 Class Syntrophia Waite et al. 2020 [monotypic]
                   Order Syntrophalesᵀ Waite et al. 2020
                     Family Syntrophaceaeᵀ Kuever et al. 2006 [monotypic]
                       1 Genus Syntrophusᵀ Mountfort et al. 1984