Superfamily Scarabaeoidea Latreille, 1802
   Family Scarabaeidae Latreille, 1802 - scarab beetles, dung beetles, and june "bugs"
     Subfamily Rutelinae MacLeay, 1819
       Tribe Rutelini MacLeay, 1819
         1 Subtribe Areodina Burmeister, 1844
         2 Subtribe Heterosternina Bates, 1888
         3 Subtribe Didrepanephorina Ohaus, 1918
         4 Subtribe Lasiocalina Ohaus, 1918
         5 Subtribe Parastasiina Burmeister, 1844
         6 Subtribe Oryctomorphina Burmeister, 1847
         7 Subtribe Desmonychina Arrow, 1917
         8 Subtribe Rutelina MacLeay, 1819