Phylum Arthropoda Siebold, 1848 - arthropods
   Class Insecta Linnaeus, 1758 - insects
     Subclass Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
       Endopterygota Sharp, 1899
         Order Lepidoptera Linnaeus, 1758 - butterflies, moths
           Suborder Heteroneura Tillyard, 1918 see Infraorder Heteroneura Tillyard, 1918
             Superfamily Papilionoidea Latreille, 1802
               Family Nymphalidae Rafinesque, 1815 - brush-footed butterflies
                 Subfamily Argynninae P.A.J. Duponchel, [1835] see Tribe Argynnini Swainson, 1833
                   Section II see Tribe Melitaeini Herrich-Schäffer, 1843
                     1 Genus Euphydryas Scudder, 1872
                     2 Genus Mellicta Billberg, 1820 see Genus Melitaea Fabricius, 1807
                     3 Genus Melitaea Fabricius, 1807
                     4 Genus Microtia Bates, 1864
                     5 Genus Chlosyne Butler, 1870
                     6 Genus Phyciodes Hübner, 1819
                     7 Genus Gnathotriche Felder & Felder, 1862
                     8 Genus Timelaea Lucas, 1883