Family Nymphalidae Rafinesque, 1815 - brush-footed butterflies
   Subfamily Nymphalinae Rafinesque, 1815
     Tribe Melitaeini Herrich-Schäffer, 1843
       Subtribe "Gnathotrichina" Kons, 2000, nom. nud.
         1 Genus Gnathotriche Felder & Felder, 1862
         2 Genus Higginsius Hemming, 1964