prokaryotes - bacteria see Domain Bacteria Woese et al. 1990
   Domain "Bacteria" Woese et al. 1990
     Phylum "Acidobacteria" Thrash and Coates 2012
       Class Holophagae Fukunaga et al. 2008
         Order Holophagales Fukunaga et al. 2008 [monotypic]
           Family Holophagaceae Fukunaga et al. 2008
             Genus Holophaga Liesack et al. 1995