Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Subphylum Chelicerata Heymons, 1901
Euchelicerata Weygoldt & Paulus, 1979
Prosomapoda Lamsdell, 2013
Planaterga Lamsdell, 2013
Dekatriata Lamsdell, 2013
Sclerophorata Kamenz, Staude & Dunlop, 2011
Class Arachnida Lamarck, 1801
Subclass Dromopoda Shultz, 1990 [paraphyletic]
Haplocnemata Börner, 1904, sedis mutabilis
Order Pseudoscorpiones De Geer, 1778
Suborder Heterosphyronida Chamberlin, 1929
Superfamily Chthonioidea Daday, 1888
Family Chthoniidaeᵀ Daday, 1888
Subfamily Chthoniinaeᵀ Daday, 1888
Tribe Chthoniiniᵀ Daday, 1888
01 Genus †Weygoldtiella Harvey, Cosgrove, Harms, Selden, Shih & Wang, 2018
02 Genus Aphrastochthonius Chamberlin, 1962
03 Genus Apochthonius Chamberlin, 1929
04 Genus Austrochthonius Chamberlin, 1929
05 Genus Caribchthonius Muchmore, 1976
06 Genus Chiliochthonius Vitali-di Castri, 1976
07 Genus Chthoniusᵀ Koch, 1843
08 Genus Congochthonius Beier, 1959
09 Genus Drepanochthonius Beier, 1964
10 Genus Francochthonius Vitali-di Castri, 1976
11 Genus Kleptochthonius Chamberlin, 1949
12 Genus Malcolmochthonius Benedict, 1978
13 Genus Mexichthonius Muchmore, 1975
14 Genus Mundochthonius Chamberlin, 1929
15 Genus Neochthonius Chamberlin, 1929
16 Genus Pseudochthonius Balzan, 1892
17 Genus Sathrochthoniella Beier, 1967
18 Genus Sathrochthonius Chamberlin, 1962
19 Genus Spelyngochthonius Beier, 1955
20 Genus Tyrannochthoniella Beier, 1966