Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Hymenopterida
Order Hymenopteraᵀ Linnaeus, 1758 - ants, bees, and wasps
Neohymenoptera
Euhymenoptera
Suborder Apocrita Gerstaecker, 1867
Aculeata Latreille, 1802
Superfamily Apoidea Latreille, 1802
Anthophilaᵀ Latreille, 1804 - bees
Family Megachilidae Latreille, 1802
Subfamily Megachilinaeᵀ Latreille, 1802
Tribe Osmiini Newman, 1834
Genus Afroheriades Peters, 1970
Genus Ashmeadiella Cockerell, 1897
Genus Atoposmia Cockerell, 1935
Genus Bekilia Benoist, 1963
Genus Chelostoma Latreille, 1809
Genus Haetosmia Popov, 1952
Genus Hofferia Tkalcu, 1984
Genus Hoplitis Klug, 1807
Genus Hoplosmia Thomson, 1872
Genus Noteriades Cockerell, 1931
Genus Ochreriades Mavromoustakis, 1956
Genus Othinosmia Michener, 1943
Genus Protosmia Ducke, 1900
Genus Pseudoheriades Peters, 1970
Genus Stenoheriades Tkalcu, 1984
Genus Stenosmia Michener, 1941
Genus Wainia Tkalcu, 1980
Genus Xeroheriades Griswold, 1986
1 Subtribe Heriadina Michener, 1941
2 Subtribe Osmiina Newman, 1834