Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Domain Bacteria Woese et al. 2024
             Kingdom Pseudomonadati (Gibbons and Murray 1978) Oren and Göker 2024
               Phylum Pseudomonadotaᵀ Garrity et al. 2021
                 Class Alphaproteobacteria Garrity et al. 2006
                   Order Hyphomicrobiales Douglas 1957 (Approved Lists 1980)
                     Family Methylocystaceae Bowman 2006
                       Genus Methylocystisᵀ (ex Whittenbury et al. 1970) Bowman et al. 1993
                         Methylocystis heyeri Dedysh et al. 2007