Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         [crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade natates Cavalier-Smith, 2021
                   Clade photaria Cavalier-Smith, 2021
                     Clade eucorta Cavalier-Smith, 2021
                       Superkingdom Corticata Lankester, 1878 - corticates
                         Kingdom Chromista Caval.-Sm. (1981) - chromists
                           Subkingdom Harosa Cavalier-Smith, 2010
                             [crown] Clade Sar Adl et al. 2012 [Simpson & Dunthorn 2020]
                               Infrakingdom Rhizaria Cavalier-Smith, 2002
                                 Phylum Cercozoa Cavalier-Smith, 1998
                                   Subphylum Reticulofilosa Cavalier-Smith, 1997
                                     Class Granofilosea D. Bass et al., 2009
                                       Order Desmothoracida Hertwig & Lesser, 1874
                                         Genus Penardiophrys K.A. Mikrjukov, 2000
                                           Penardiophrys reticulataᵀ (Penard, 1904) K.A. Mikrjukov, 2000
                                           Penardiophrys spinifera (Brown, 1918) K.A. Mikrjukov, 2000