Clade †Pterosauria Owen 1842 [Andres & Padian 2020] see Order Pterosauria Kaup, 1834
   Clade †Macronychoptera Unwin 2003 [Andres 2021]
     Clade †Novialoidea Kellner 2003 [Andres 2021]
       Clade †Breviquartossa Unwin 2003 [Andres 2021]
         Clade †Rhamphorhynchidae Hoernes 1884 [Andres 2021] see Family Rhamphorhynchidae Seeley, 1870
           Clade †Rhamphorhynchinae Nopcsa 1928 [Andres 2021] see Subfamily Rhamphorhynchinae Seeley, 1870
             Clade †Rhamphorhynchae Seeley 1870 [Andres 2021]
               Clade †Angustinaripterini Andres, Clark & Xu 2014 [Andres 2021] see Tribe Angustinaripterini He, Yan & Su, 1983
                 Genus †Harpactognathus Carpenter, Unwin, Cloward, Miles & Miles, 2003