Superfamily Scarabaeoidea Latreille, 1802
   Family Geotrupidae Latreille, 1802
     Subfamily Bolboceratinae Mulsant, 1842
       1 Tribe Bolboceratini Mulsant, 1842
       2 Tribe Athyreini Howden & Martínez, 1963
       2.0 Group Athyréites Blanchard, 1845, nom. inval. see Tribe Athyreini Howden & Martínez, 1963
       3 Tribe Bolbochromini Nikolajev, 1970
       4 Tribe Gilletinini Nikolajev, 1990
       5 Tribe Eubolbitini Nikolajev, 1970
       6 Tribe Bolbelasmini Nikolajev, 1996
       7 Tribe Australobolbini Nikolajev, 1996
       8 Tribe Eucanthini Nikolajev, 2002b
       9 Tribe Stenaspidiini Nikolajev, 2003