Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Adephaga Schellenberg, 1806
Family Carabidae Latreille, 1802 - ground beetles
Subfamily Pterostichinae
[Genus Europhilus Chaudoir, 1859]
see Subgenus Europhilus {Chaudoir, 1859}
[Genus Laemosthenes auct., Schaufuss, 1865]
see Genus Pristonychus Dejean, 1828
01 Genus Abacidus Leconte, 1863
02 Genus Abax Bonelli, 1810
02.0 [Genus "Abas" Ernst, van Andel, Drent, Tinbergen & Bakker, 1985, orth. inval.]
see Genus Abax Bonelli, 1810
03 Genus Agonum Bonelli, 1810 ex Stephens, 1827
04 Genus Antisphodrus Schaufuss, 1865
05 Genus Calathus Bonelli, 1810 ex Stephens, 1827
06 Genus Dolichus Bonelli, 1809
07 Genus Dysidius Chaudoir, 1838
08 Genus Euferonia Casey, 1918
09 Genus Eumolops Casey, 1918
10 Genus Gastrellarius Casey, 1918
11 Genus Idiochroma Bedel, 1902
12 Genus Melanius Dejean, 1828
13 Genus Molops Bonelli, 1810
14 Genus "Odontonyx" auct.
15 Genus Olisthopus Dejean, 1828
16 Genus Platyderus Stephens, 1828
17 Genus Platynus Stephens, 1827
18 Genus Poecilus Samouelle, 1819
19.1 Genus Pristonychus Dejean, 1828
19.2 Genus Laemostenus Bonelli, 1810 ex Agassiz, 1846
19.2.0 [Genus "Laemosthenes" Bonelli, 1810, nom. nud.]
see Genus Laemosthenes Bonelli, 1810 ex Agassiz, 1846
19.2.0 [Genus Laemosthenes Bonelli, 1810 ex Agassiz, 1846]
see Genus Laemostenus Bonelli, 1810 ex Agassiz, 1846
20 Genus Pseudargutor Casey, 1918
21 Genus Pterostichusᵀ Stephens, 1827
22 Genus Sphodrus Clairville, 1806
23 Genus Stomis Clairville, 1806
24 Genus Synuchus Gyllenhal, 1810