[viruses]
   Realm Riboviria
     Order Nidovirales
       Suborder Cornidovirineae [monotypic]
         Family Coronaviridae
           Subfamily Orthocoronavirinae
             Genus Betacoronavirus
               Subgenus Merbecovirus
                 1 Hedgehog coronavirus 1
                 2 Middle East respiratory syndrome-related coronavirus
                 3 Pipistrellus bat coronavirus HKU5
                 4 Tylonycteris bat coronavirus HKU4