Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Adephaga Schellenberg, 1806
Family Carabidae Latreille, 1802 - ground beetles
Subfamily Callistinae Habe & Kosuge, 1967
Genus Chlaenius Dejean, 1826
Subgenus Chlaenius (Agostenus) Motschulsky, 1850
Subgenus Chlaenius (Chlaeniellus) {Reitter, 1908}
Subgenus Chlaenius (Chlaenites) {Motschulsky, 1860}
Subgenus Chlaenius (Chlaenius)ᵀ Bonelli, 1809
Chlaenius aestivus Say, 1823
Chlaenius cordicollis Kirby, 1837
Chlaenius diffinis Chaudoir, 1856
Chlaenius erythropus Germar, 1824
Chlaenius impunctifrons Say, 1823
Chlaenius leucoscelis Chevrolat, 1834
Chlaenius nebraskensis LeConte, 1856
Chlaenius nemoralis Say, 1823
Chlaenius pennsylvanicus Say, 1823
Chlaenius propeagilis Liu et al., 2011
Chlaenius rotundithorax Liu et al., 2010
Chlaenius sericeus (Forster, 1771)
Chlaenius tomentosus (Say, 1823)
Chlaenius tricolor Dejean, 1826
Subgenus Chlaenius (Dinodes) {Bonelli, 1809}
Subgenus Chlaenius (Pelasmus) {Motschulsky, 1850}