Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Cynopterus Cuvier, 1824 - short-nosed fruit bats
           Cynopterus titthaecheilus (Temminck, 1825)
             Cynopterus titthaecheilus terminus Sody, 1940