Lepidoptera Linnaeus, 1758 - butterflies, moths
   Superfamily Hesperioidea Latreille, 1809 [monotypic] see Superfamily Papilionoidea Latreille, 1802
     Family Hesperiidae Latreille, 1809 - skipper butterflies
       Subfamily Pyrginae Burmeister, 1878
         Tribe Pyrgini Burmeister, 1878
           Genus group Telemiades Group
             Genus subgroup Quadrus Subgroup
               1 Genus Gorgythion Godman & Salvin, 1896
               2 Genus Ouleus Lindsey, 1925
               3 Genus Zera Evans, 1953
               4 Genus Quadrus Lindsey, 1925