 Natura - nature
 Natura - nature
   actualia - actual entities
 actualia - actual entities
     Mundus Plinius - physical world
 Mundus Plinius - physical world
       naturalia - natural bodies
 naturalia - natural bodies
         Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
 Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
 Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
 Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
 Clade discaria Cavalier-Smith, 2021
                 Clade dorsates Cavalier-Smith, 2021
 Clade dorsates Cavalier-Smith, 2021
                   Clade podiates Cavalier-Smith, 2012
 Clade podiates Cavalier-Smith, 2012
                     Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
 Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
                       Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
 Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
                         Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
 Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
                           Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
 Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                             Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
 Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
                               Kingdom Animalia Linnaeus, 1758 - animals
 Kingdom Animalia Linnaeus, 1758 - animals
                                 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                   Subkingdom Bilateria Hatschek, 1888 - bilaterians
 Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                     Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
 Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                       Infrakingdom Protostomia Grobben, 1908
 Infrakingdom Protostomia Grobben, 1908
                                         Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
 Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                           Panarthropoda Nielsen, 1995 - panarthropods
 Panarthropoda Nielsen, 1995 - panarthropods
                                             Tactopoda Budd, 2001
 Tactopoda Budd, 2001
                                               Phylum Arthropoda Siebold, 1848 - arthropods
 Phylum Arthropoda Siebold, 1848 - arthropods
                                                 Euarthropoda
 Euarthropoda
                                                   Antennulata
 Antennulata
                                                     Subphylum Mandibulata Snodgrass, 1938
 Subphylum Mandibulata Snodgrass, 1938
                                                       Infraphylum Pancrustacea Zrzavý & Štys, 1997
 Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                         Altocrustacea
 Altocrustacea
                                                           Miracrustacea
 Miracrustacea
                                                             Superclass Hexapoda Latreille, 1825
 Superclass Hexapoda Latreille, 1825
                                                               Cercophora
 Cercophora
                                                                 Class Insecta Linnaeus, 1758 - insects
 Class Insecta Linnaeus, 1758 - insects
                                                                   Subclass Dicondylia Haeckel, 1866
 Subclass Dicondylia Haeckel, 1866
                                                                     Infraclass Pterygota Lang, 1888
 Infraclass Pterygota Lang, 1888
                                                                       Metapterygota
 Metapterygota
                                                                         Neoptera Martynov, 1923
 Neoptera Martynov, 1923
                                                                           Eumetabola
 Eumetabola
                                                                             Holometabola
 Holometabola
                                                                               Superorder Hymenopterida
 Superorder Hymenopterida
                                                                                 Order Hymenopteraᵀ Linnaeus, 1758 - ants, bees, and wasps
 Order Hymenopteraᵀ Linnaeus, 1758 - ants, bees, and wasps
                                                                                   Neohymenoptera
 Neohymenoptera
                                                                                     Euhymenoptera
 Euhymenoptera
                                                                                       Suborder Apocrita Gerstaecker, 1867
 Suborder Apocrita Gerstaecker, 1867
                                                                                         Aculeata Latreille, 1802
 Aculeata Latreille, 1802
                                                                                           Superfamily Vespoidea Latreille, 1802
 Superfamily Vespoidea Latreille, 1802
                                                                                             Family Formicidae Latreille, 1809 - ants
 Family Formicidae Latreille, 1809 - ants
                                                                                               Subfamily group "myrmicomorph group"
 Subfamily group "myrmicomorph group"
                                                                                                 Subfamily Myrmicinaeᵀ Lepeletier de Saint Fargeau, 1835
 Subfamily Myrmicinaeᵀ Lepeletier de Saint Fargeau, 1835
                                                                                                   Group "formicoxenite tribal group"
 Group "formicoxenite tribal group"
                                                                                                     Tribe Formicoxeniniᵀ Forel, 1893
 Tribe Formicoxeniniᵀ Forel, 1893
                                                                                                       Genus Leptothorax Mayr, 1855
 Genus Leptothorax Mayr, 1855
                                                                                                         01 Leptothorax acervorum (Fabricius, 1793)
 01 Leptothorax acervorum (Fabricius, 1793)
                                                                                                         02 Leptothorax athabasca Buschinger & Schulz
 02 Leptothorax athabasca Buschinger & Schulz
                                                                                                         03 Leptothorax buschingeri Kutter, 1967
 03 Leptothorax buschingeri Kutter, 1967
                                                                                                         04 Leptothorax calderoni Creighton
 04 Leptothorax calderoni Creighton
                                                                                                         05 Leptothorax crassipilis Wheeler, 1917
 05 Leptothorax crassipilis Wheeler, 1917
                                                                                                         06 Leptothorax faberi Buschinger, 1983
 06 Leptothorax faberi Buschinger, 1983
                                                                                                         07.1 Leptothorax goesswaldi Kutter, 1967
 07.1 Leptothorax goesswaldi Kutter, 1967
                                                                                                         07.2 Leptothorax kutteri Buschinger, 1966
 07.2 Leptothorax kutteri Buschinger, 1966
                                                                                                         07.3 Leptothorax pacis (Kutter, 1945)
 07.3 Leptothorax pacis (Kutter, 1945)
                                                                                                         07.4 Leptothorax pocahontas (Buschinger, 1979)
 07.4 Leptothorax pocahontas (Buschinger, 1979)
                                                                                                         08 Leptothorax gredleri Mayr, 1855
 08 Leptothorax gredleri Mayr, 1855
                                                                                                         09 Leptothorax muscorum (Nylander, 1846)
 09 Leptothorax muscorum (Nylander, 1846)
                                                                                                         10 Leptothorax oceanicus (Kuznetsov-Ugamsky, 1928)
 10 Leptothorax oceanicus (Kuznetsov-Ugamsky, 1928)
                                                                                                         11 Leptothorax palustris Cover & Deyrup
 11 Leptothorax palustris Cover & Deyrup
                                                                                                         12 Leptothorax paraxenus Heinze & Alloway, 1992
 12 Leptothorax paraxenus Heinze & Alloway, 1992
                                                                                                         13 Leptothorax retractus Francoeur, 1986
 13 Leptothorax retractus Francoeur, 1986
                                                                                                         14 Leptothorax scamni Ruzsky, 1905
 14 Leptothorax scamni Ruzsky, 1905
                                                                                                         15 Leptothorax sphagnicola Francoeur, 1986
 15 Leptothorax sphagnicola Francoeur, 1986
                                                                                                         16 Leptothorax wilsoni Heinze, 1989
 16 Leptothorax wilsoni Heinze, 1989