Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Hymenopterida
Order Hymenopteraᵀ Linnaeus, 1758 - ants, bees, and wasps
Neohymenoptera
Euhymenoptera
Suborder Apocrita Gerstaecker, 1867
Aculeata Latreille, 1802
Superfamily Vespoidea Latreille, 1802
Family Vespidaeᵀ Latreille, 1802
Genus Ancistrocerus Wesmael, 1836
Genus Brachygastra Perty, 1833
Genus Cephalodynerus Parker, 1965
Genus Dolichodynerus Bohart, 1939
Genus Eumenes Latreille, 1802
Genus Euodynerus Dalla Torre, 1904
Genus Euparagia Cresson, 1879
Genus Gastrodynerus Bohart, 1984
Genus Hypalastoroides Saussure, 1856
Genus Leptochiloides Bohart, 1940
Genus Leptochilus Saussure, 1853
Genus Leucodynerus Bohart, 1982
Genus Maricopodynerus Viereck, 1908
Genus Microdynerus Thomson, 1874
Genus Mischocyttarus Saussure, 1853
Genus Monobia Saussure, 1852
Genus Montezumia Saussure, 1852
Genus Odynerus Latreille, 1803
Genus "Odynerus" auct.
Genus Pachodynerus Saussure, 1870
Genus Parancistrocerus Bequaert, 1925
Genus Paranortonia Bequaert, 1940
Genus Polistes auct.
Genus Polybia Lepeletier, 1836
Genus Pseudepipona Saussure, 1856
Genus Pseudodynerus Saussure, 1855
Genus Pseudomasaris Ashmead, 1902
Genus Pterocheilus Klug, 1805
Genus Smeringodynerus Snelling, 1975
Genus Stenodynerus Saussure, 1863
Genus Symmorphus Wesmael, 1836
Genus Zethus Fabricius, 1804
1 Subfamily Stenogastrinae
2 Subfamily Polistinae (Lepeletier de Saint-Fargeau, 1835)
3 Subfamily Vespinaeᵀ Latreille, 1802