Phylum Arthropoda Siebold, 1848 - arthropods
   Class Insecta Linnaeus, 1758 - insects
     Subclass Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
       Endopterygota Sharp, 1899
         Order Lepidoptera Linnaeus, 1758 - butterflies, moths
           Suborder Heteroneura Tillyard, 1918 see Infraorder Heteroneura Tillyard, 1918
             Superfamily Papilionoidea Latreille, 1802
               Family Nymphalidae Rafinesque, 1815 - brush-footed butterflies
                 Subfamily Charaxinae Guenée, 1865
                   1 Tribe Preponini Rydon, 1971
                   2 Tribe Prothoini Roepke, 1938
                   3 Tribe Charaxini Guenée, 1865
                   4 Tribe Anaeini Reuter, 1896