Phylum Arthropoda Siebold, 1848 - arthropods
   Class Insecta Linnaeus, 1758 - insects
     Subclass Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
       Endopterygota Sharp, 1899
         Order Lepidoptera Linnaeus, 1758 - butterflies, moths
           Suborder Heteroneura Tillyard, 1918 see Infraorder Heteroneura Tillyard, 1918
             Superfamily Papilionoidea Latreille, 1802
               Family Satyridae Boisduval, 1833 see Subfamily Satyrinae Boisduval, 1833
                 Subfamily Elymniinae see Tribe Elymniini Herrich-Schäffer, 1864
                   Tribe Zetherini Reuter, 1896
                     1 Genus Zethera Felder, 1861
                     2 Genus Callarge Leech, 1892
                     3 ?Genus Penthema Doubleday, 1848