Phylum Mollusca Linnaeus, 1758 - molluscs
   Class Bivalvia Linnaeus, 1758 - bivalves
     Subclass Palaeoheterodonta Newell, 1965 see Clade Palaeoheterodonta Newell, 1965
       Order Unionida Gray, 1854
         Superfamily Unionoidea Rafinesque, 1820
           Family Unionidae Rafinesque, 1820 - pearly mussels
             Subfamily Ambleminae Rafinesque, 1820
               Tribe Amblemini Rafinesque, 1820
                 Genus Quadrula Rafinesque, 1820
                   1 Subgenus Amphinaias Fischer & Crosse, 1894
                   2 Subgenus Luteacarnea Thiele, 1934
                   3 Subgenus Obliquata Frierson, 1927
                   4 Subgenus Pleuronaia Frierson, 1927