Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus niger species group
             1 Pteropus aldabrensis True, 1893
             2 Pteropus niger (Kerr, 1792) - greater Mascarene flying fox
             3 Pteropus rufus Geoffroy, 1803 - Madagascar flying fox
             4 Pteropus seychellensis Milne-Edwards, 1877 - Seychelles flying fox
             5 Pteropus voeltzkowi Matschie, 1909 - Pemba flying fox