Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus vampyrus species group
             1 Pteropus giganteus (Brünnich, 1782) - Indian flying fox
             2 Pteropus intermedius Andersen, 1908
             3 Pteropus lylei Andersen, 1908 - Lyle's flying fox
             4 Pteropus vampyrus (Linnaeus, 1758) - large flying fox