Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus scapulatus species group
             1 Pteropus gilliardorum Van Deusen, 1969
             2 Pteropus mahaganus Sanborn, 1931 - lesser flying fox
             3 Pteropus scapulatus Peters, 1862 - little red flying fox
             4 Pteropus woodfordi Thomas, 1888 - least flying fox