Clade †Pterosauria Owen 1842 [Andres & Padian 2020] see Order Pterosauria Kaup, 1834
   Clade †Macronychoptera Unwin 2003 [Andres 2021]
     Clade †Novialoidea Kellner 2003 [Andres 2021]
       Clade †Breviquartossa Unwin 2003 [Andres 2021]
         Clade †Pterodactylomorpha Andres, Clark & Xu 2014 [Andres 2021] see Suborder Pterodactylomorpha Schaller, 2007
           Clade †Monofenestrata Lü, Unwin, Jin, Liu & Ji 2010 [Andres 2021]
             Clade †Darwinoptera Andres, Clark & Xu 2014 [Andres 2021]
               Clade †Wukongopteridae Wang, Kellner, Jiang & Meng 2009 [Andres 2021] see Family Wukongopteridae Wang, Kellner, Jiang & Meng, 2009
                 Clade †Wukongopterinae Zhou, Pêgas, Ma, Han, Jin, Leal, Bonde, Kobayashi, Lautenschlager, Wei, Shen & Ji. 2021
                   Genus †Darwinopterus Lü, Unwin, Jin, Liu & Ji, 2009