†Pterosauria Owen 1842 [Sereno 1991], nom. cl. inval. see Clade Pterosauria Owen 1842 [Andres & Padian 2020]
   †Novialoideaᵀ Kellner 2003 nom. cl. inval. [within unnamed taxon] see Clade Novialoidea Kellner 2003 [Andres 2021]
     †Pterodactyloideaᵀ Plieninger 1901 [Kellner 2003], nom. cl. inval. [within unnamed taxon] see Clade Pterodactyloidea Plieninger 1901 [Andres & Padian 2020]
       †Dsungaripteroideaᵀ Young 1964 [Kellner 2003] see Superfamily Dsungaripteroidea Young, 1964
         †Nyctosauridaeᵀ Hay 1902 [Kellner 2003] nom. cl. inval. see Clade Nyctosauridae Hay 1902 [Andres 2021]
           Genus †Nyctosaurusᵀ Marsh, 1876
             1 †Nyctosaurus gracilis (Marsh, 1876)
             2 †Nyctosaurus bonneri (Miller, 1971)
             3 †'Nyctosaurus' lamegoi Price, 1953