Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus subniger species group
             Pteropus dasymallus Temminck, 1825 - Ryukyu flying fox
               1 [Pteropus dasymallus dasymallus] Temminck, 1825
               2 Pteropus dasymallus daitonensis Kuroda, 1921
               3 Pteropus dasymallus formosus Sclater, 1873
               4 Pteropus dasymallus inopinatus Kuroda, 1933
               5 Pteropus dasymallus yayeyamae Kuroda, 1933