Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus subniger species group
             Pteropus griseus (Geoffroy, 1810) - grey flying fox
               1 [Pteropus griseus griseus] (Geoffroy, 1810)
               2 Pteropus griseus mimus Andersen, 1908
               3 Pteropus griseus pallidus Temminck, 1825