Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus mariannus species group
             Pteropus tonganus Quoy & Gaimard, 1830 - insular flying fox
               1 [Pteropus tonganus tonganus] Quoy & Gaimard, 1830
               2 Pteropus tonganus basiliscus Thomas, 1915
               3 Pteropus tonganus geddiei MacGillivray, 1860