Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Rousettus Gray, 1821 - rousette fruit bats
           1 Subgenus Rousettus Gray, 1821
           2 Subgenus Boneia Jentink, 1879
           3 Subgenus Stenonycteris Gray, 1871