Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Rhinolophidae Gray, 1825 - horseshoe bats
         Genus Rhinolophus Lacépède, 1799 - horseshoe bats
           Rhinolophus pusillus species group
             Rhinolophus cornutus Temminck, 1834 - little Japanese horseshoe bat
               1 [Rhinolophus cornutus cornutus] Temminck, 1834
               2 Rhinolophus cornutus miyakonis Kuroda, 1924
               3 Rhinolophus cornutus orii Kuroda, 1924
               4 Rhinolophus cornutus perditus Andersen, 1918
               5 Rhinolophus cornutus pumilus Andersen, 1905