Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Hipposideridae Lydekker, 1891 - Old World leaf-nosed bats
         Genus Hipposideros Gray, 1831 - Old World leaf-nosed bats
           Hipposideros bicolor species group
             Hipposideros ater Templeton, 1848 - dusky leaf-nosed bat
               1 [Hipposideros ater ater] Templeton, 1848
               2 Hipposideros ater amboinensis Peters, 1871
               3 Hipposideros ater antricola Peters, 1861
               4 Hipposideros ater aruensis Gray, 1858
               5 Hipposideros ater gilberti Johnson, 1959
               6 Hipposideros ater nicobarulae Miller, 1902
               7 Hipposideros ater saevus Andersen, 1918