living organisms see Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
   Superkingdom Eukaryotae see Domain Eukaryota Chatton, 1925
     Kingdom Animalia Linnaeus, 1758 - animals
       Subkingdom Eumetazoa Butschli, 1910 see [unranked] Eumetazoa Bütschli, 1910
         Phylum Arthropoda Siebold, 1848 - arthropods
           Subphylum Uniramia Snodgrass, 1935 see Phylum Arthropoda Siebold, 1848
             Class Insecta Linnaeus, 1758 - insects
               Subclass Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
                 Infraclass Neoptera Martynov, 1923 - modern, wing-folding insects see Division Neoptera Martynov, 1923
                   Superorder Holometabola - holometabolans see Subdivision Endopterygota Sharp, 1899
                     Order Coleoptera Linnaeus, 1758 - beetles
                       Suborder Polyphaga Emery, 1886
                         Superfamily Scarabaeoidea Latreille, 1802
                           Family Geotrupidae Latreille, 1802
                             1 Subfamily Geotrupinae Latreille, 1802
                             2 Subfamily Lethrinae Mulsant & Rey, 1871 see Tribe Lethrini Mulsant & Rey, 1871
                             3 Subfamily Taurocerastinae Germain, 1897
                             4 Subfamily Athyreinae Howden & Martínez, 1963 see Tribe Athyreini Howden & Martínez, 1963
                             5 Subfamily Bolboceratinae Mulsant, 1842