Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
[crown] Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
[crown] Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
[crown] Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Hymenopterida
Order Hymenopteraᵀ Linnaeus, 1758 - ants, bees, and wasps
Neohymenoptera
Euhymenoptera
Suborder Apocrita Gerstaecker, 1867
Aculeata Latreille, 1802
Superfamily Chrysidoidea Latreille, 1802
Family Bethylidae Forster, 1856
01 Genus Anisepyris Kieffer, 1906
02 Genus Apenesia Westwood, 1874
03 Genus Bakeriella Kieffer, 1910
04 Genus Cephalonomia Westwood, 1833
05 Genus Dissomphalus Ashmead, 1893
06 Genus Glenosema Kieffer, 1905
07 Genus Goniozus Förster, 1856
08 Genus Holepyris Kieffer, 1904
09 Genus Laelius Ashmead, 1893
10 Genus Nesepyris Bridwell, 1920
11 Genus Parascleroderma Kieffer, 1904
12 Genus Plastanoxus Kieffer, 1905
13 Genus Prorops Waterston, 1923
14 Genus Prosierola Förster, 1905
15 Genus Pseudisobrachium Kieffer, 1904
16 Genus Rhabdepyris Kieffer, 1904
17 Genus Sclerodermus Latreille, 1809
18 Genus Sierola Cameron, 1881
19 Subfamily Bethylinaeᵀ Halliday, 1840
20 Subfamily Pristocerinae Mocsary, 1881
21 Subfamily Epyrinae Kieffer, 1914
22 Subfamily Mesitiinae (Kieffer, 1914)