Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           "neomura" Cavalier-Smith 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               neokaryotes Cavalier-Smith 1993 - mitochondrial eukaryotes
                 neozoa Cavalier-Smith 1993
                   scotokaryotes Cavalier-Smith 2000
                     podiates Cavalier-Smith 2012 [nom. cl. nov.]
                       Amorphea Adl et al. 2012 [nom. cl. nov.]
                         Opisthokonta Cavalier-Smith 1987
                           Superkingdom Holomycota Liu, Steenkamp, Brinkmann, Forget, Philippe & Lang, 2009
                             1 Kingdom Nucleariae Tedersoo, Sánchez-Ramírez, Kõljalg, Bahram, Döring, Schigel, May, Ryberg & Abarenkov, 2018 [monotypic]
                             2 Kingdom Fungi R.T. Moore (1980) - fungi