Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
[crown] Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade natates Cavalier-Smith, 2021
Clade photaria Cavalier-Smith, 2021
Clade eucorta Cavalier-Smith, 2021
Superkingdom Corticata Lankester, 1878 - corticates
Kingdom Plantae Haeckel (1866) - plants
Subkingdom Viridiplantae Caval.-Sm. (1981) - green plants
Infrakingdom Streptophyta Caval.-Sm. (1993)
[crown] Clade Phragmoplastophyta Lecointre & Le Guyader 2006 [Hall, McCourt, Delwiche, Mishler & Karol 2020]
[crown] Clade Embryophytaᵀ Engler 1887 [Mishler & Qiu 2020]
Phylum Bryophytaᵀ A.Braun (1864) - bryophytes
[crown] Clade "Setaphyta" Puttick, Morris, Williams, Cox, Edwards, Kenrick, Pressel, Wellman, Schneider, Pisani & Donoghue 2018
Subphylum Marchantiophytina Doweld (2001) - liverworts
Class Jungermanniopsida Stotler & Crand.-Stotl. (1977) - leafy liverworts
Subclass Jungermanniidaeᵀ Engl. (1893)
Order Jungermannialesᵀ H. Klinggr.
Suborder Lophocoleineae Schljakov
1 Family Pseudolepicoleaceae Fulford & J. Taylor
2 Family Trichocoleaceae Nakai
3 Family Grolleaceae Solari ex R.M. Schust. [monotypic]
4 Family Mastigophoraceae R.M. Schust.
5 Family Herbertaceae Müll. Frib. ex Fulford & Hatcher
6.1 Family Vetaformaceae Fulford & J.Taylor (1963) [monotypic]
6.2 Family Lepicoleaceae R.M. Schust. [monotypic]
7.1 Family Phycolepidoziaceae R.M. Schust. [monotypic]
7.2 Family Lepidoziaceae Limpr.
8.1 Family Lophocoleaceaeᵀ Vanden Berghen
8.2 Family Brevianthaceae J.J. Engel & R.M.Schust. (1981) [monotypic]
8.3 Family Chonecoleaceae R.M. Schust. ex Grolle [monotypic]
8.4 Family Plagiochilaceae Müll. Frib. & Herzog